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GMD | Articles | Volume 12, issue 8
Geosci. Model Dev., 12, 3649–3685, 2019
© Author(s) 2019. This work is distributed under
the Creative Commons Attribution 4.0 License.

Special issue: Paleoclimate Modelling Intercomparison Project phase 4 (PMIP4)...

Geosci. Model Dev., 12, 3649–3685, 2019
© Author(s) 2019. This work is distributed under
the Creative Commons Attribution 4.0 License.

Model experiment description paper 22 Aug 2019

Model experiment description paper | 22 Aug 2019

The penultimate deglaciation: protocol for Paleoclimate Modelling Intercomparison Project (PMIP) phase 4 transient numerical simulations between 140 and 127 ka, version 1.0

The penultimate deglaciation: protocol for Paleoclimate Modelling Intercomparison Project (PMIP) phase 4 transient numerical simulations between 140 and 127 ka, version 1.0
Laurie Menviel1, Emilie Capron2,3, Aline Govin4, Andrea Dutton5, Lev Tarasov6, Ayako Abe-Ouchi7, Russell N. Drysdale8,9, Philip L. Gibbard10, Lauren Gregoire11, Feng He12, Ruza F. Ivanovic11, Masa Kageyama4, Kenji Kawamura13,14,15, Amaelle Landais4, Bette L. Otto-Bliesner16, Ikumi Oyabu13, Polychronis C. Tzedakis17, Eric Wolff18, and Xu Zhang19,20 Laurie Menviel et al.
  • 1Climate Change Research Center, PANGEA, the University of New South Wales, Sydney, Australia
  • 2Physics of Ice, Climate and Earth, Niels Bohr Institute, University of Copenhagen, Tagensvej 8, 2100 Copenhagen, Denmark
  • 3British Antarctic Survey, High Cross, Madingley Road, Cambridge, CB3 0ET, UK
  • 4Laboratoire des Sciences du Climat et de l'Environnement (LSCE), Institut Pierre Simon Laplace (IPSL), CEA-CNRS-UVSQ, Université Paris-Saclay, Gif-Sur-Yvette, 91190, France
  • 5Department of Geological Sciences, University of Florida, P.O. Box 112120, Gainesville, FL 32611, USA
  • 6Department of Physics and Physical Oceanography, Memorial University of Newfoundland, St John's, Canada
  • 7Atmosphere and Ocean Research Institute, The University of Tokyo, Tokyo, Japan
  • 8School of Geography, The University of Melbourne, Melbourne, Australia
  • 9Laboratoire EDYTEM UMR CNRS 5204, Université Savoie Mont Blanc, 73376 Le Bourget du Lac, France
  • 10Scott Polar Research Institute, University of Cambridge, Cambridge, CB2 1ER, UK
  • 11School of Earth and Environment, University of Leeds, Leeds, LS2 9JT, UK
  • 12Center for Climatic Research, Nelson Institute for Environmental Studies, University of Wisconsin-Madison, Madison, WI 53706, USA
  • 13National Institute of Polar Research, Research Organizations of Information and Systems, 10-3 Midori-cho, Tachikawa, Tokyo 190-8518, Japan
  • 14Department of Polar Science, Graduate University for Advanced Studies (SOKENDAI), 10-3 Midori-cho, Tachikawa, Tokyo 190-8518, Japan
  • 15Institute of Biogeosciences, Japan Agency for Marine-Earth Science and Technology, 2-15 Natsushima-cho, Yokosuka 237-0061, Japan
  • 16Climate and Global Dynamics Laboratory, National Center for Atmospheric Research (NCAR), Boulder, CO 80305, USA
  • 17Environmental Change Research Centre, Department of Geography, University College London, London, UK
  • 18Department of Earth Sciences, University of Cambridge, Cambridge, CB2 3EQ, UK
  • 19Alfred Wegener Institute, Helmholtz Centre for Polar and Marine Research, 27570 Bremerhaven, Germany
  • 20Key Laboratory of Western China's Environmental Systems (Ministry of Education), College of Earth and Environmental Sciences, Lanzhou University, Lanzhou, 730000, China

Correspondence: Laurie Menviel ( and Emilie Capron (

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The penultimate deglaciation (PDG, ∼138–128 thousand years before present, hereafter ka) is the transition from the penultimate glacial maximum (PGM) to the Last Interglacial (LIG, ∼129–116 ka). The LIG stands out as one of the warmest interglacials of the last 800 000 years (hereafter kyr), with high-latitude temperature warmer than today and global sea level likely higher by at least 6 m. Considering the transient nature of the Earth system, the LIG climate and ice-sheet evolution were certainly influenced by the changes occurring during the penultimate deglaciation. It is thus important to investigate, with coupled atmosphere–ocean general circulation models (AOGCMs), the climate and environmental response to the large changes in boundary conditions (i.e. orbital configuration, atmospheric greenhouse gas concentrations, ice-sheet geometry and associated meltwater fluxes) occurring during the penultimate deglaciation.

A deglaciation working group has recently been set up as part of the Paleoclimate Modelling Intercomparison Project (PMIP) phase 4, with a protocol to perform transient simulations of the last deglaciation (19–11 ka; although the protocol covers 26–0 ka). Similar to the last deglaciation, the disintegration of continental ice sheets during the penultimate deglaciation led to significant changes in the oceanic circulation during Heinrich Stadial 11 (∼136–129 ka). However, the two deglaciations bear significant differences in magnitude and temporal evolution of climate and environmental changes.

Here, as part of the Past Global Changes (PAGES)-PMIP working group on Quaternary interglacials (QUIGS), we propose a protocol to perform transient simulations of the penultimate deglaciation under the auspices of PMIP4. This design includes time-varying changes in orbital forcing, greenhouse gas concentrations, continental ice sheets as well as freshwater input from the disintegration of continental ice sheets. This experiment is designed for AOGCMs to assess the coupled response of the climate system to all forcings. Additional sensitivity experiments are proposed to evaluate the response to each forcing. Finally, a selection of paleo-records representing different parts of the climate system is presented, providing an appropriate benchmark for upcoming model–data comparisons across the penultimate deglaciation.

1 Introduction
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Over the last 450 kyr, Earth's climate has been dominated by glacial–interglacial cycles with a recurrence period of about 100 kyr. These asymmetrical cycles are characterized by long glacial periods, associated with a gradual cooling, a slow decrease in atmospheric greenhouse gas (GHG) concentrations and a progressive growth of large continental ice sheets in the Northern Hemisphere (NH), leading to a 60 to 120 m global sea-level decrease (Lisiecki and Raymo2005; Grant et al.2014; Rohling et al.2017). These long glacial periods were followed by relatively rapid multi-millennial-scale warmings into consecutive interglacial states. These deglaciations represent the largest natural global warming and large-scale climate reorganizations across the Quaternary. Deglaciations are paced by an external forcing, i.e. variations in the seasonal and latitudinal distribution of incoming solar radiation (insolation) driven by changes in Earth's orbit (Berger1978). However, changes in insolation alone are not sufficient to explain the amplitude of these major warmings and require amplification mechanisms. These amplification mechanisms are related to the large increase in atmospheric GHG concentrations (e.g. atmospheric CO2 increases by 60 to 100 ppm; Lüthi et al.2008), the disintegration of NH ice sheets and their associated change in albedo (Abe-Ouchi et al.2013), as well as changes in sea-ice and vegetation cover (Fig. 1). Hence, deglaciations provide a great opportunity to study the interaction between the different components of the Earth System and climate's sensitivity to changes in radiative forcing.

Figure 1Overview of (left) the penultimate and (right) the last deglaciations climatic and environmental evolutions: (a) Hovmöller diagram of summer solstice insolation anomalies (W m−2). This corresponds to 21 June in the Northern Hemisphere and 21 December in the Southern Hemisphere. Time series of (b) eccentricity (red), obliquity (solid black) and precession (dashed black) (Berger1978). (c) Global mean sea-level anomaly probability maximum (m, blue), (left) including its 95 % confidence interval (cyan) (Grant et al.2014), from Grant et al. (2012) for the original age model for the penultimate and (right) from Lambeck et al. (2014) for the last deglaciation. (d) Atmospheric methane (CH4) concentration as recorded in the EDC ice core, Antarctica (Loulergue et al.2008). (e) Precipitation-weighted surface temperature reconstruction based on stable water isotopes from the Greenland NEEM ice core (left) and annual surface temperature composite reconstruction based on air nitrogen isotopes from the Greenland NEEM, NGRIP and GISP2 ice cores (Buizert et al.2014). (f) Alkenone-based (Uk'37) SST reconstruction from ODP976 (Martrat et al.2014). (g) Atmospheric CO2 concentration as recorded in Antarctic ice cores for (left) EDC (Bereiter et al.2015) and (right) WAIS divide(Marcott et al.2014) on the WD2014 chronology (Buizert et al.2015). (h) Antarctic temperature anomalies relative to the present day inferred from the EDC ice core (Jouzel et al.2007). Unless specified differently, all records are displayed on the AICC2012 timescale (Bazin et al.2013; Veres et al.2013) or a chronology coherent with AICC2012.


A pervasive characteristic of the five deglaciations of the past 450 kyr is the occurrence of millennial-scale climate events (e.g. Cheng et al.2009; Barker et al.2011; Vázquez Riveiros et al.2013; Past Interglacials Working Group of PAGES2016; Rodrigues et al.2017). In the North Atlantic, these events, also referred to as stadials, are characterized by a substantial weakening of North Atlantic Deep Water (NADW) formation (e.g. McManus et al.2004; Vázquez Riveiros et al.2013; Böhm et al.2015; Ng et al.2018), possibly due to meltwater discharge into the North Atlantic (Ivanovic et al.2018). There is a link between these events and enhanced iceberg calving, supported by the concurrent presence of ice-rafted debris (IRD) in North Atlantic marine sediment cores, and the stadials that contain substantial IRD layers (identified as Heinrich events) are known as Heinrich stadials (e.g. Heinrich1988; Bond et al.1992; McManus et al.1999; van Kreveld et al.2000; Hodell et al.2017).

A weakening of the Atlantic meridional heat transport during these stadials maintains cold conditions at high northern latitudes in the Atlantic sector (Stouffer et al.2007; Swingedouw et al.2009; Kageyama et al.2010, 2013) while contributing to a gradual warming at high southern latitudes (Blunier and Brook2001; Stocker and Johnsen2003; EPICA community members2006), thus leading to a bipolar seesaw pattern of climate changes. The strengthening of NADW formation at the end of stadials induces a relatively abrupt temperature increase in the northern North Atlantic and surrounding regions and a sharp increase in atmospheric CH4 and Asian monsoon strength (Loulergue et al.2008; Cheng et al.2009; Buizert et al.2014). While a significant atmospheric CO2 increase is also observed during the NADW recovery at the end of Heinrich stadials during deglaciations (Marcott et al.2014), the major phase of atmospheric CO2 increase coincides with a Southern Ocean (e.g. Barker et al.2009; Uemura et al.2018) and Antarctic warming (Cheng et al.2009; Masson-Delmotte et al.2011; Landais et al.2013; Marcott et al.2014) (Fig. 1). The sequence of events leading to the deglacial atmospheric CO2 increase is still poorly constrained. Still, it most likely resulted from a combination of processes (e.g. Kohfeld and Ridgwell2009), including changes in solubility, global alkalinity content (e.g. Sigman et al.2010), iron fertilization (e.g. Martin1990; Bopp et al.2003; Martínez-García et al.2014), Antarctic sea-ice cover (Stephens and Keeling2000) and ocean circulation (e.g. Toggweiler et al.2006; Anderson et al.2009; Skinner et al.2010; Toggweiler and Lea2010). Changes in ocean circulation, and particularly variations in the formation rates of the main deep and bottom water masses, i.e. NADW and Antarctic Bottom Water, can significantly impact atmospheric CO2 by modifying the vertical gradient in oceanic dissolved inorganic carbon (e.g. Menviel et al.2014, 2017, 2018).

While they share similarities, the last two deglaciations also bear significant differences in amplitude and durations. Figure 1 shows the evolution of key variables across 15 kyr, from glacial maxima to peak interglacial conditions. The two deglaciations initiate under a range of glacial ice-sheet states and progress under a variety of orbital-forcing scenarios (Tzedakis et al.2017; Past Interglacials Working Group of PAGES2016) (Fig. 1). Although there are still many open questions, the sequence of events occurring during the last deglaciation, which represents the transition from the Last Glacial Maximum (LGM; Marine Isotope Stage 2, hereafter MIS2, 26–19 ka) to our current interglacial, is starting to emerge (Cheng et al.2009; Denton et al.2010; Shakun et al.2012). The last deglaciation began with Heinrich Stadial 1 (HS1, ∼ 18–14.7 ka) when part of the Laurentide and Eurasian ice sheets disintegrated (Dyke2004; Hughes et al.2016), draining freshwater to the North Atlantic, that may have contributed to the observed weakening of NADW formation (e.g. McManus et al.2004; Gherardi et al.2009; Thornalley et al.2011; Ng et al.2018; Ivanovic et al.2018). HS1 was characterized by cold and dry conditions in the North Atlantic, over Greenland and Europe (Tzedakis et al.2013; Buizert et al.2014; Martrat et al.2014). Antarctic temperature and global atmospheric CO2 concentration rose during this period. Paleo-proxy records and modelling studies suggest that the Intertropical Convergence Zone (ITCZ) shifted southward (e.g. Arz et al.1998; Chiang and Bitz2005; Timmermann et al.2005; Stouffer et al.2007; Kageyama et al.2009, 2013; Marzin et al.2013; McGee et al.2014), thus leading to dry conditions in most of the northern tropics, including the northern part of South America (Peterson et al.2000; Deplazes et al.2013; Montade et al.2015) and the Sahel (Mulitza et al.2008; Niedermeyer et al.2009). Chinese speleothem records also indicate a weak East Asian summer monsoon activity during HS1 (Wang et al.2001; Cheng et al.2009).

The abrupt NADW resumption at ∼14.7 ka, imprinted by a sharp atmospheric CH4 concentration increase (Loulergue et al.2008), led to a warm period in the North Atlantic region, the Bølling–Allerød (Liu et al.2009; Buizert et al.2014) (Fig. 1, right). This period of North Atlantic warming coincides with (and may have triggered part of) the period of fastest sea-level rise, Meltwater Pulse-1A (MWP-1A) (Gregoire et al.2016), during which sea level rose by 14–18 m in less than 500 years, starting at ∼14.5 ka (e.g. Deschamps et al.2012; Lambeck et al.2014). At high southern latitudes, the gradual deglacial warming was interrupted by the Antarctic Cold Reversal (ACR, ∼14.5–12.8 ka) (Jouzel et al.1995, 2007; Pedro et al.2016), which was also coincident with a pause in the deglacial atmospheric CO2 increase (e.g. Marcott et al.2014). The ACR could be the result of enhanced northward heat transport from the Southern Hemisphere due to the strong NADW resumption occurring during the Bølling–Allerød (Pedro et al.2016) or from a meltwater pulse originating from the Antarctic ice sheet at the time of MWP-1A (Weaver et al.2003; Menviel et al.2011; Weber et al.2014; Golledge et al.2014).

A return to stadial conditions over Greenland, Europe and the North Atlantic occurred during the Younger Dryas (∼12.8–11.7 ka; Fig. 1, right) (Alley2000). This event likely resulted from a combination of processes (Renssen et al.2015), possibly including a weakening of NADW formation resulting from an increase in meltwater discharge into the Arctic Ocean (Tarasov and Peltier2005; Murton et al.2010; Keigwin et al.2018) or melting of the Fennoscandian ice sheet (Muschitiello et al.2015), and an altered atmospheric circulation due to a minimum in solar activity (Renssen et al.2000). While the Barbados coral record suggests that a second phase of rapid sea-level rise occurred at about 11.3 ka (MWP-1B) (e.g. Bard et al.1990), data from Tahiti boreholes (Bard et al.2010) and from a compilation of sea-level data (Lambeck et al.2014) provide no evidence of a particularly rapid sea-level rise during that period.

The penultimate deglaciation (∼138–128 ka, referred to here as PDG), which represents the transition between the penultimate glacial maximum (PGM) (MIS 6, also referred to as Late Saalian, 160–140 ka) and the Last Interglacial (LIG; also referred to as MIS 5e in marine sediment cores) (Govin et al.2015), has received less attention. The PGM was characterized by an atmospheric CO2 content of ∼195 ppm (Lüthi et al.2008) and a significantly different extent of NH ice sheets compared to the LGM (Dyke et al.2002; Svendsen et al.2004; Lambeck et al.2006; Ehlers et al.2011; Margari et al.2014). The eustatic sea level during the PGM is estimated at ∼90–100 m lower than present-day values (Rabineau et al.2006; Grant et al.2012; Rohling et al.2017), with a relatively large uncertainty range (Rohling et al.2017). This compares to ∼130 m lower or more than during the present day during the LGM (Austermann et al.2013; Lambeck et al.2014). The LIG also bears significant differences to the interstadial that followed the last deglaciation, i.e. the Holocene. Latest data-based estimates suggest that sea level was ∼6 to 9 m higher during the LIG than today, thus implying a significant ice-mass loss from both the Greenland and Antarctic ice sheets (e.g. Dutton et al.2015). In addition, compared to pre-industrial times, high-latitude sea surface temperature (SST) and Greenland surface temperatures were respectively ≥1C and 3 to 11 C greater during the LIG (e.g. Landais et al.2016; Capron et al.2017; Hoffman et al.2017). Considering the transient nature of the Earth system, a better understanding of the PDG could thus improve our knowledge of the processes that led to continental ice-mass loss during the LIG.

Recent work (Masson-Delmotte et al.2011; Landais et al.2013; Govin et al.2015) also depicted a sequence of events over the PDG that contrasts with the one across the last deglaciation. Paleo-proxy records indicate that the disintegration of NH ice sheets induced a ∼80 m sea-level rise (Grant et al.2014) between 135 and 129 ka. This is also concomitant with Heinrich Stadial 11 (HS11) (e.g. Heinrich1988; Oppo et al.2006; Skinner and Shackleton2006; Govin et al.2015). HS11 was characterized by weak NADW formation (Oppo et al.2006; Böhm et al.2015; Deaney et al.2017), cold and dry conditions in the North Atlantic region (Drysdale et al.2009; Martrat et al.2014; Marino et al.2015), and gradually warmer conditions over Antarctica (Jouzel et al.2007), associated with a sustained atmospheric CO2 increase of ∼60 ppm between ∼134 and 129 ka (Landais et al.2013) (Figs. 1, 2). Increasing evidence of sub-millennial-scale climate changes at high and low latitudes during HS11 (e.g. Martrat et al.2014) prompts the need to refine the sequence of events across the PDG. However, this is challenging as (i) climatic reconstructions over the PDG are still scarce and most records have insufficient resolution to allow the identification of centennial- to millennial-scale climatic variability and (ii) it is difficult to establish robust absolute and relative chronologies for most paleo-climatic records across this time interval (Govin et al.2015).

While our knowledge of the processes and feedbacks occurring during deglaciations has significantly improved over the last two decades (e.g. Cheng et al.2009, 2016; Shakun et al.2012; Abe-Ouchi et al.2013; Landais et al.2013), many unknowns remain. For example, our understanding of the precise roles of atmospheric and oceanic processes in leading to the waning of glacial continental ice sheets during deglaciations is still incomplete. It is also crucial to comprehend the subsequent impacts of continental ice-sheet disintegration on the oceanic circulation, climate, the terrestrial vegetation and the carbon cycle system.

Numerical simulations performed with climate models provide a dynamical framework to understand the response of the Earth system to external forcing (i.e. insolation) and internal dynamics (e.g. albedo, GHGs) that culminate in deglaciations. Atmospheric and oceanic teleconnections associated with millennial-scale variability can also be studied in detail. Model–paleo-climate proxy comparisons, including snapshot experiments at 130 and at 126 ka, suggest that the inclusion of freshwater forcing in the North Atlantic due to the melting of NH ice sheets could explain the relatively cold conditions in the North Atlantic and warm conditions in the Southern Ocean observed in paleo-data during these time periods (Govin et al.2012; Stone et al.2016). However, snapshot experiments assume that the climate state is in near-equilibrium, and because relatively rapid and large changes in both internal and external forcings occur during deglaciations, transient simulations (i.e. numerical simulations with time-varying boundary conditions) are needed. These simulations also allow a more robust paleo-data–modelling comparison, thus enabling the refinement of the sequence of events.

Transient numerical simulations have already been performed for the last deglaciation (Liu et al.2009; Menviel et al.2011; Roche et al.2011; Gregoire et al.2012; He et al.2013; Otto-Bliesner et al.2014) and provide a dynamical framework to further our understanding of the climate-change drivers, teleconnections and feedbacks inherent in the Earth system. Currently, no three-dimensional transient simulations were run across the full time interval covered by the penultimate deglaciation (∼140–127 ka). However, transient simulations covering the period 135 to 115 ka have been performed with a range of models to understand the impact of surface boundary conditions and freshwater fluxes on the LIG (Bakker et al.2013; Loutre et al.2014; Goelzer et al.2016a). The need for transient simulations is now recognized, and a protocol to perform transient experiments of the last deglaciation as part of Paleoclimate Modelling Intercomparison Project (PMIP) phase 4 has been recently established (Ivanovic et al.2016). However, to further our understanding of the processes at play during deglaciations, including the role of millennial-scale climate change, other deglaciations should be studied in detail. Transient simulations of the penultimate deglaciation could also help to understand the climate and sea-level highstand occurring during the LIG. We thus propose to extend the PMIP4 working group on the last deglaciation to include the penultimate deglaciation and thus create a deglaciation working group. This effort will complement the last deglaciation experiments of PMIP4 (Ivanovic et al.2016), allowing an evaluation of the similarities and differences in the climate system response during the last and the penultimate deglaciations.

Here we present a protocol to perform transient numerical simulations of the PDG from 140 to 127 ka with coupled atmosphere–ocean general circulation models (AOGCMs). These experiments will provide a link with the PMIP4 transient LIG experiment (127 to 121 ka) (Otto-Bliesner et al.2017). After a description of changes in insolation (Sect. 2), GHGs (Sect. 3), continental ice sheets (Sect. 4) and sea level (Sect. 5) occurring during the PDG, we present a framework to perform transient simulations of PDG (Sects. 6 and 7), as well as a selection of key paleo-climate and paleo-environmental records to be used for model–data comparisons (Sect. 8).

Figure 2Atmospheric greenhouse gas concentrations: atmospheric trace gases through the penultimate deglaciation from Antarctic ice cores displayed on the AICC2012 chronology (Bazin et al.2013; Veres et al.2013) and the spline that should be used to force the transient simulations (solid and dotted black lines; Köhler et al.2017a). (a) Atmospheric CO2 concentrations from EDC (turquoise and blue) (Lourantou et al.2010; Schneider et al.2013) and TALDICE (green) (Schneider et al.2013). (b) Atmospheric CH4 concentration from EDC (Loulergue et al.2008) (blue), Vostok (Petit et al.1999) (orange), TALDICE (Buiron et al.2011) (green) and EDML (Capron et al.2010) (pink). (c) Atmospheric N2O concentration from EDC (Flückiger et al.2002) (blue), EDML (Schilt et al.2010) (pink) and TALDICE (Schilt et al.2010) (green). Due to in situ production within the ice sheet, no accurate N2O measurements are available beyond 134.5 ka. Between 140 and 134.5 ka N2O should increase linearly from 201 to 218.74 ppb (dashed black line). Red crosses indicate the 140 ka spin-up values for CO2, CH4 and N2O concentrations (i.e. 191 ppm, 385 and 201 ppb, respectively).


2 Insolation
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The orbital parameters (eccentricity, obliquity and longitude of perihelion) should be time evolving and set following Berger (1978). This external forcing affects the seasonal and latitudinal distribution, as well as the magnitude of solar energy received at the top of the atmosphere and, in the case of obliquity (Earth's axial tilt), the annual mean insolation at any given latitude with opposite, but small, effects at low and high latitudes. Eccentricity is high during the entire deglaciation period, ranging from 0.033 at 140 ka to 0.041 at 127 ka, and is significantly higher than during the last deglaciation (Fig. 1b; ∼0.019 at the LGM to 0.020 at 14 ka) and than the present value of 0.0167. Obliquity peaks at 131 ka; the degree of tilt is similar between the last and the penultimate deglaciation. Perihelion occurs near the NH winter solstice at 140 ka, shifting to near the NH summer solstice by 127 ka.

Although the overall trends in summer solstice insolation anomalies, as compared to the mean of the last 1000 years, evolve similarly in the last and penultimate deglaciations, the magnitudes of the maximum positive summer anomalies in the NH and the minimum negative summer anomalies in the Southern Hemisphere are much greater during the PDG, when eccentricity is higher, than during the last deglaciation (Fig. 1a). At 65 N, peak summer anomalies of more than 70 W m−2 occur at 128 ka. In contrast, during the last deglaciation, the 65 N summer solstice anomalies peak at 11 ka, with anomalies of ∼50 W m−2. Similarly, 65 S summer solstice negative anomalies are close to −40 W m−2 at 127 ka but only about −20 W m−2 at 10 ka. In addition, the rates of change in summer solstice anomalies are greater from 140 to 127 ka than from 21 to 8 ka.

Given the clear differences between the solar forcing of last and penultimate deglaciations, comparing the two transient deglacial simulations will provide valuable information on the underlying mechanisms and Earth system feedbacks. The solar constant should be set to 1360.7 W m−2, consistent with the Coupled Model Intercomparison Project phase 6 (CMIP6)–PMIP4 piControl and lig127k simulations (Otto-Bliesner et al.2017) as well as the PMIP4 transient climate simulation of the last deglaciation (Ivanovic et al.2016).

3 Greenhouse gases
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GHG records are available solely from Antarctic ice cores across the time interval 140–127 ka (Fig. 2). LIG GHG records from North Greenland Eemian Ice Drilling (NEEM) and other Greenland ice cores are affected by stratigraphic disturbances and in situ CO2, CH4 and N2O production (e.g. Tschumi and Stauffer2000; NEEM community members2013). The North Greenland Ice Project (NGRIP) ice core provides a continuous and reliable CH4 record, but it only extends back to ∼123 ka (North Greenland Ice Core project members2004). We first briefly describe existing atmospheric CO2, CH4 and N2O records (below). The recent spline-smoothed GHG curves calculated from a selection of those records (Köhler et al.2017a) should be used. They have the benefit of providing continuous GHG records, with a temporal resolution of 1 year on the commonly used AICC2012 gas-age scale (Bazin et al.2013; Veres et al.2013). This timescale is associated with an average 1σ absolute error of ∼2 kyr between 140 and 127 ka.

Atmospheric CO2 concentrations have been measured on the EPICA Dome C (EDC) and Talos Dome Ice core (TALDICE) ice cores (Fig. 2). The EDC records from Lourantou et al. (2010) and Schneider et al. (2013) agree well overall. The Schneider et al. (2013) dataset depicts a long-term CO2 increase starting at ∼137.8 ka and ending at ∼128.5 ka with a centennial-scale CO2 rise above the subsequent LIG CO2 values, also referred to as an “overshoot”. The CO2 overshoot is smaller in the Schneider et al. (2013) dataset compared to a similar feature measured in Lourantou et al. (2010): while the former displays a relatively constant CO2 concentration of ∼275 ppm between 128 and 126 ka, the latter shows a CO2 decrease from 280 to 265 ppm between 128 and 126 ka. The offsets between CO2 records from the same EDC core are likely related to the different air extraction techniques used in the two studies (Schneider et al.2013). The smoothed spline CO2 curve, which should be used as forcing for the PDG is based on those two EDC datasets and the calculation method accounts for such potential difference in local maxima (details provided in Köhler et al.2017a).

Atmospheric CH4 concentration records from Vostok, EPICA Dronning Maud Land (EDML), EDC and TALDICE agree well within the gas-age uncertainties attached to each core (Fig. 2). They illustrate a slow rise from ∼390 to 540 ppb between ∼137 and 129 ka, which is followed by an abrupt increase of ∼200 ppb reaching maximum LIG values at ∼128.5 ka. Because CH4 sources are located mostly in the NH, an interpolar concentration difference (IPD) between Greenland and Antarctic CH4 records exists.

For instance, an IPD of ∼14, ∼34 and ∼43 ppb is reported during the LGM, Heinrich Stadial 1 and the Bølling warming respectively (Baumgartner et al.2012). However, without reliable CH4 records from Greenland ice cores, it remains challenging to estimate the evolution of the IPD across the deglaciation. Hence, for the atmospheric CH4 forcing of transient PDG simulations, the smoothed spline CH4 curve, which is solely based on the EDC CH4 record (Köhler et al.2017a) should be used, recognizing that the values may be 1 %–4 % lower than the actual global average.

Both CO2 and CH4 concentrations undergo some rapid changes around 140 ka, which is also the time when the models should spin up. To avoid possible artificial abrupt changes in the GHG, the average values obtained for the interval 139–141 (i.e. 191 ppm for CO2 and 385 ppb for CH4, Table 1) should be used as spin-up CO2 and CH4 concentrations. Consequently, CO2 and CH4 changes between 140 and 139 ka provided in the forcing scenarios are linearly interpolated between the 140 ka spin-up values and those at 139 ka of 196.68 ppm for CO2 and 287.65 ppb for CH4. From 139 ka, the spline-smoothed curves from Köhler et al. (2017a) should be used.

Berger (1978)Berger (1978)Berger (1978)(Bazin et al.2013; Veres et al.2013)Köhler et al. (2017a)(Lourantou et al.2010; Schneider et al.2013)Köhler et al. (2017a)(Loulergue et al.2008)Köhler et al. (2017a)(Schilt et al.2010; Spahni et al.2005)(Abe-Ouchi et al.2013)(Tarasov et al.2012)(Briggs et al.2014)

Table 1Summary of forcings and boundary conditions to apply for the PGM spin-up (140 ka) and subsequent transient simulation of the PDG.

Download Print Version | Download XLSX

Atmospheric TALDICE, EDML and EDC N2O records are available between 134.5 and 127 ka (Fig. 2) (Schilt et al.2010; Flückiger et al.2002). From 134.5 to 128 ka, N2O levels increase from ∼220 to 270 ppb. Following a short decrease until ∼127 ka, N2O concentrations stabilize afterwards. No reliable atmospheric N2O concentrations are available beyond 134 ka as N2O concentrations measured in the air trapped in ice from the PGM are affected by in situ production related to microbial activity (Schilt et al.2010). During the LGM (considered here as the time interval 26–21 ka), the average N2O level was ∼201 ppb. Assuming the LGM is an analogue for the PGM, we propose a 140 ka spin-up value and N2O transient forcing curve that starts with a 201 ppb level and then linearly increases to 218.74 ppb at 134.5 ka. From 134.5 ka, the N2O smoothed spline curve calculated by Köhler et al. (2017a), which is based on the TALDICE and EDC discrete N2O measurements, should be used.

The CO2 and N2O levels from the spline curves at 127 ka (274 ppm and 257 ppb) only differ from the values chosen as boundary conditions for the PMIP4 lig127k equilibrium experiment by 1 ppm and 2 ppb respectively (Otto-Bliesner et al.2017; Köhler et al.2017a). The comparison is less direct for CH4. Indeed, a global CH4 value (685 ppm) rather than an Antarctic ice-core-based CH4 value (e.g. CH4 level of 660 ppm at 127 ka in Köhler et al.2017a) is proposed as forcing for the lig127k simulations. However, this difference in global atmospheric CH4 and Antarctic ice-core CH4 concentration is similar to the one observed during the mid-Holocene (23 ppb) (Otto-Bliesner et al.2017; Köhler et al.2017a).

4 Continental ice sheets
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Changes in continental ice sheets during the deglaciation will significantly impact the climate system through their albedo, which will directly affect the radiative balance (e.g. He et al.2013). Changes in continental ice-sheet geometry can also significantly impact atmospheric dynamics (e.g. Zhang et al.2014; Gong et al.2015). Transient simulations of the PDG will thus need to be forced by the three-dimensional and time-varying evolution of continental ice sheets, which is currently only available from numerical simulations. However, simulating the evolution of continental ice sheets across the PDG is associated with large uncertainties, due to the climate forcing of the ice-sheet models and poorly constrained non-linearities within the ice-sheet system. Glacial geological data (e.g. glacial deposits and glacial striations) are also available to constrain continental ice-sheet evolutions and can thus provide an estimate of the uncertainties associated with the numerical ice-sheet evolutions. In this section, we describe the available numerical ice-sheet evolutions to use as a forcing of the transient simulations of the PDG. We further compare the results of these simulations with existing glacial geological constraints.

4.1 Combined ice-sheet forcing

To facilitate the transient simulations of the PDG, we provide a combined ice-sheet forcing (available in the Supplement and on the PMIP4 wiki), in which separate reconstructions of different ice sheets have been merged. As the sea-level solver assumes an equilibrium initial condition, the simulations start at the previous interglacial. As is standard, the solver also requires present-day ice-sheet histories to bias-correct against present-day observed topography. Thus, a full 240 kyr ice-sheet history is required. The simulated NH ice-sheet evolution, described in Sect. 4.2 (Abe-Ouchi et al.2013), is merged with the simulated Greenland and the Antarctic (Briggs et al.2014) evolutions described in Sect. 4.3 and 4.4, respectively. The resolution of the merged ice-sheet file is 1 longitude by 0.5 latitude. The merger involves no extra smoothing, beyond that inherent in the glacial isostatic adjustment solver, which involves transformation to spherical harmonics. The merger involves a simple masking operation with the mask boundary through Nares Strait, Baffin Bay, Davis Strait and the Labrador Sea. Examination of the resultant topography shows small merger artefacts around Nares Strait ranging to a few hundred metres in elevation difference.

From the LIG onward, the combined ice-sheet evolution, referred to as GLAC-1D in PMIP4, is used (e.g. Ivanovic et al.2016). GLAC-1D includes the Greenland and Antarctic ice-sheet components described in Sect. 4.3 and 4.4 (Briggs et al.2014), the North American ice-sheet simulation described in Tarasov et al. (2012), and the Eurasian ice-sheet simulation described in Tarasov (2014). The ice-sheet thickness from these simulations is run through a sea-level solver using the Viscosity Model (VM5a) (Peltier and Drummond2008) Earth rheology to extract a gravitationally self-consistent topography. The surface topography is then run through a global surface drainage solver (using the algorithm described in Tarasov and Peltier2006) to extract the relevant surface drainage pointer field for each time slice. This will indicate into which ocean grid cell each terrestrial grid cell will drain.

4.2 North American and Eurasian ice sheets

The evolution of NH ice sheets during the PDG is given by a numerical simulation performed with the thermo-mechanically coupled ice-sheet model IcIES (Ice-sheet model for Integrated Earth system Studies) with an original resolution of 1 by 1 in the horizontal and 26 vertical levels (Abe-Ouchi et al.2007) (Fig. 3). IcIES uses the shallow ice approximation and computes the evolution of grounded ice but not floating ice shelves. The sliding velocity is related to the gravitational driving stress according to Payne (1999) and basal sliding only occurs when the basal ice is at the pressure melting point. This ice-sheet model was driven by climatic changes obtained from the MIROC general circulation model (GCM) (Abe-Ouchi et al.2013), which was forced by changes in insolation and atmospheric CO2 concentration. In global agreement with glacial geological constraints (Dyke et al.2002; Svendsen et al.2004; Curry et al.2011; Syverson and Colgan2011) and other numerical simulations of NH ice-sheet evolution (Tarasov et al.2012; Abe-Ouchi et al.2013; Peltier et al.2015; Colleoni et al.2016), the simulated extent and volume of the North American ice sheet was smaller during the PGM than the LGM (Fig. 3).

Figure 3NH ice-sheet elevation (m) at (a) 140, (b) 136, (c) 134, (d) 132, (e) 130 and (f) 128 ka from the combined ice-sheet forcing as simulated by the IcIES-MIROC model (Abe-Ouchi et al.2013) for the North American and Eurasian ice sheets and as simulated by the Glacial Systems Model (GSM) (e.g. Tarasov et al.2012) for the Greenland ice sheet. Elevation is shown where the ice mask is greater than 0.5.

In Eurasia, the PGM recorded the most extensive glaciation since MIS 12 (Hughes and Gibbard2019). The maximum extent of the Fennoscandian ice sheet probably occurred at ∼160 ka, when it extended into the central Netherlands, Germany and the Russian Plain (Margari et al.2010; Ehlers et al.2011; Hughes and Gibbard2019). This was followed by a partial melting of the Fennoscandian ice sheet, peaking between ∼157 and 154 ka, and a readvance after 150 ka (Margari et al.2010; Hughes and Gibbard2019). The maximum extent of the NH ice sheets probably occurred at the end of the PGM (Margari et al.2014; Head and Gibbard2015). In Europe the late PGM ice advance was less expensive than at ∼160 ka but that was compensated for by ice-sheet expansion in Russia, Siberia (Astakhov et al.2016) and North America (e.g. Curry et al.2011; Syverson and Colgan2011). Glacial geological constraints (e.g. Astakhov2004; Svendsen et al.2004) indeed suggest that the Barents–Kara ice sheet extended further during the PGM than the LGM. The simulated Eurasian ice sheet is in general agreement with the reconstruction of Lambeck et al. (2006), with a dome reaching 3000 m over the Kara Sea during the PGM that subsequently disintegrated across the deglaciation. However, the extent and volume of the simulated Eurasian ice sheet might be underestimated since it is smaller at the PGM than LGM, whereas reconstructions suggest it should be larger at the PGM (Lambeck et al.2006; Rohling et al.2017).

Rohling et al. (2017) further suggest that the ice volume was almost equally distributed between Eurasia and North America at the PGM, with a 33 to 53 m global mean sea-level equivalent (sle) contribution from the Eurasian ice sheet and 39–59 m from North America, whereas the ice-sheet simulation produces a ∼24 m sle contribution from Eurasia. Thus, the volume of the North American ice sheet may also be overestimated.

In the ice-sheet simulation, NH ice-mass loss closely follows the boreal summer insolation and occurs mostly between ∼134 and 127 ka (Fig. 4), with two peaks of glacial meltwater release at ∼131 and 128 ka. By 132 ka, the Eurasian ice sheet has decreased significantly and the southern and western flanks of the North American ice sheet have disintegrated (Fig. 3). Another significant retreat of the North American ice sheet occurs between 132 and 128 ka, at which point it is mostly restricted to the north of the Hudson Bay. By 127 ka, the North American ice sheet only remains over Baffin Island.

Figure 4Time series of (a) summer solstice insolation at 65 N and 65 S, (b)(d) global sea-level equivalent (m) of changes in (b) NH (Abe-Ouchi et al.2013), (c) Greenland (Tarasov et al.2012) and (d) Antarctic ice sheets (Briggs et al.2014), (e) and global sea-level (m) change estimated from Red Sea records (Grant et al.2014) (green) with the 95 % probability interval (cyan), sea-level record with an adjusted age scale as described in Sect. 5 (blue) and as estimated from the continental ice-sheet simulations (black). The red line shows the changes in global sea level that would be obtained by adding the meltwater flux, fIRD, described in (f) plus an Antarctic contribution of 13 m. (f) Possible North Atlantic meltwater flux (Sv) scenarios: estimated from the disintegration of NH ice sheets as shown in (b, c) (Tarasov et al.2012; Abe-Ouchi et al.2013) (black), estimated from the global sea-level change on the revised age scale (this study, blue) and scaled from the North Atlantic and Norwegian Sea IRD records (red, shown in Fig. 8b).


4.3 Greenland ice sheet

The Greenland model uses an updated version (GSM.G7.31.18) of the Glacial Systems Model (e.g. Tarasov et al.2012) run at grid resolution of 0.5 longitude by 0.25 latitude. The model has been upgraded to hybrid shallow-ice and shallow-shelf physics, with an ice dynamical core from Pollard and DeConto (2012) and includes a 4 km deep permafrost-resolving bed thermal component (Tarasov and Peltier2007), a viscoelastic bedrock response with global ice-sheet and sea-level loading, sub-shelf melt, parametrizations for subgrid mass-balance and ice flow (Le Morzadec et al.2015), and updated parametrizations for surface mass-balance and ice calving.

Given that with active bed thermodynamics (down to 4 km), the thermodynamic equilibration timescale is greater than 100 kyr for the Greenland ice sheet, the most appropriate method is to start the run during the previous interglacial period. Therefore, the model runs start at 240 ka with present-day ice and bedrock geometry and with an ice and bed temperature field from the end of a previous 240 ka model run. The model is then forced from 240 ka until 0 ka, with a climate forcing that is partly glacial-index-based, using a composite of a glaciological inversion of the GISP2 regional temperature change (for the last 40 kyr) and the synthetic Greenland δ18O curve that was deduced from the Antarctic EDC isotopic record assuming a thermal bipolar seesaw pattern (Barker et al.2011). The climate forcing also includes a two-way coupled 2-D energy balance climate model (Tarasov and Peltier1997) to capture radiative changes.

Greenland ice-sheet model runs are scored against a large set of constraints including relative sea-level (RSL), proximity to present-day ice-surface topography, present-day observed basal temperatures from various ice cores, time of deglaciation of Nares Strait and the location of the present-day summit. The last 20 kyr of the run is critical as this represents the time period with most of the data constraints for Greenland. The simulation presented here (G9175) is a least misfit model from a preliminary exploratory ensemble.

This simulation suggests no significant change in Greenland ice mass until ∼134 ka (Fig. 4), followed by a small ice-mass loss, mostly from floating ice, between 134 and 130 ka. In this simulation, the main phase of Greenland deglaciation occurs between 130 and 127 ka, during which Greenland loses an ice mass of 2.9 m sle in excess of the total pre-industrial value and then an additional 1.5 m sle. As shown in Fig. 5, the extent and height of the Greenland ice sheet is significantly smaller at 128 ka than at 132 ka. Greenland ice-mass loss is particularly evident on its western side, with a part of southwestern Greenland being ice-free. To a first order, the simulated disintegration of the Greenland ice sheet follows the increase in boreal summer insolation and in atmospheric CO2 (Fig. 1).

Figure 5Greenland ice-sheet elevation (m) at (a) 140 ka, (b) 130 ka, (c) 129 ka and (d) 128 ka as simulated by the GSM (e.g. Tarasov et al.2012). The 0 elevation contour is in black.

The main phase of the Greenland ice-sheet retreat in this simulation is globally in agreement with proxy records, which suggest significant runoff in the Labrador Sea at ∼130 ka and at ∼127 ka (e.g. Carlson and Winsor2012). However the simulated Greenland ice-sheet disintegration could be too rapid as paleo-proxy records suggest significant meltwater discharge from the Greenland ice sheet throughout the LIG (e.g. Carlson and Winsor2012). In addition, other model simulations suggest a maximum sea-level contribution from Greenland at ∼123–121 ka (Yau et al.2016; Bradley et al.2018), in agreement with the timing of the LIG minimum elevation at the Greenland NEEM location. This minimum elevation estimate was reconstructed from total air content and water isotopic records measured on the deep ice core drilled at that site (NEEM community members2013). Based on the paleo-proxy records and model simulations, the protocol for the PMIP4 LIG simulation for 127 ka (lig127k) recommends a pre-industrial Greenland configuration.

4.4 Antarctic ice sheet

The Antarctic model configuration is largely that of Briggs et al. (2013, 2014): a hybrid of the Penn State University ice-sheet model (Pollard and DeConto2012) and the GSM. Simulations are run with a 40 km grid resolution using the LR04 benthic δ18O stack (Lisiecki and Raymo2005, referred to as LR04) for sea-level forcing. The climate forcing is a parametric mix of an index-based approach (using the EDC δD record of Jouzel et al.2007) and one based on orbital forcing, as detailed by Briggs et al. (2013). The parameter vector (nn4041 from Briggs et al.2014) that gave the best fit to constraints (GSM-A) in the large ensemble analysis is used. Two changes are imposed on the model to partially rectify an inadequate LIG sea-level contribution. First, SST dependence is added to the sub-shelf melt model. Second, to compensate for inadequate LIG warming, where SSTs are above present-day values, they are then given a minimum value of 3 C (i.e. SST = MAX(SST, 3.0 C)). Even so, the Antarctic contribution to the LIG highstand is only 1.4 m sle and is therefore inadequate given current inferences (as well as constraints on contributions from Greenland and steric effects) (e.g. Kopp et al.2009).

The simulation suggests a continuous Antarctic ice-sheet discharge during the PDG, with a glacial ice-mass loss of ∼12.5 m sle between 140 and 131 ka, followed by an additional 1.4 m sle between 131 and 130 ka (Fig. 4). In this simulation, the West Antarctic ice sheet loses significant ice mass between 140 and 136 ka (Fig. 6), with a retreat of the grounding line over the Ross Sea as well as ice-mass loss in the Weddell Sea, on the Antarctic Peninsula and in the Amundsen Sea sector. By 132 ka, the grounding line has completely retreated over the Ross Sea and has retreated significantly over the Weddell Sea.

Figure 6Antarctic ice-sheet elevation at (a) 140 ka, (b) 136 ka, (c) 134 ka, (d) 132 ka, (e) 130 ka and (f) 128 ka as simulated by the GSM (e.g. Briggs et al.2014). Grounding lines are in black.

5 Sea level
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Direct evidence for constraining the evolution of the global sea level during the time interval 140–127 ka remains sparse. Although the LR04 benthic δ18O stack (Lisiecki and Raymo2005) is sometimes used to approximate sea-level change on glacial–interglacial timescales, in the case of the PDG, the timing of the LR04 benthic δ18O stack is fixed by reference to a handful of U-series coral dates from Huon Peninsula with relatively high analytical uncertainties and questionable preservation (Bard et al.1990; Stein et al.1993). Tying the MIS 5e peak to the average age of these coral dates results in a benthic δ18O minimum that is roughly centred on the main phase of coral growth during this interglacial period (122–129 ka) (Stirling et al.1998) rather than having the onset of the interglacial aligned with the timing of the onset of the sea-level highstand at far-field sites (∼129 ka) (e.g. Stirling et al.1998; Dutton et al.2015).

Here, we seek to provide an improved reconstruction of sea level across the PDG by examining available RSL records. Information on the timing and magnitude of the changes across this time interval is provided by three RSL records (Fig. 7):

  • (i)

    An RSL record from the Red Sea (Grant et al.2012), that is deduced from the planktic foraminifera δ18O measured on sediment cores retrieved in this evaporative marginal sea. This record is transformed into an RSL signal by using hydraulic models that constrain the salinity of surface waters as a function of sea level. The Red Sea record provides the only continuous profile of RSL across our interval of interest;

  • (ii)

    RSL data from the U-series dates and elevations of the submerged coral reefs of Tahiti (Thomas et al.2009); and

  • (iii)

    RSL data derived from U-series dates and elevations of uplifted coral terraces of Huon Peninsula, Papua New Guinea (Esat et al.1999).

Providing a robust age model for sediment records from the PGM to the LIG is not straightforward (e.g. Govin et al.2015), and over time, several age models have been proposed for the Red Sea RSL record (e.g. Siddall et al.2003; Rohling et al.2009; Grant et al.2012). The latest chronology is based on climatic alignment of the Red Sea RSL record to eastern Mediterranean planktic foraminifera δ18O records, which are in turn aligned to the absolutely dated Soreq Cave speleothem δ18O record (Grant et al.2012). While the absolute ages of the speleothem record have the potential to provide a more robust age model (both in terms of accuracy and precision), the application of these dates to the Red Sea sea-level reconstruction hinges on the assumption that the tie points between the Red Sea and eastern Mediterranean records have been correctly assigned, and that the intervals between these tie points can be linearly extrapolated.

The Tahiti and Huon Peninsula corals are associated with absolute radiometric dates (using U-series geochronology). For the purpose of this study, all of the U-series ages have been recalculated to normalize them with the same set of decay constants for 234U and 230Th (Cheng et al.2013) (Tables S3 and S4), using the methodology described by Hibbert et al. (2016). The array of data from Huon Peninsula suggests post-depositional alteration (open-system behaviour of the U-series isotopes) that complicates a precise age interpretation (Fig. 7).

The Red Sea time series published by Grant et al. (2012) shows that, after an RSL low stand of about −100 m relative to present between 145 and 141 ka, a brief pulse of at least ∼25 m sea-level rise, based on the smoothed record (or up to ∼50 m based on the unsmoothed time series), occurred between ∼141 and 138 ka (identified as MWP-2A in Marino et al.2015; Fig. 7a). This pulse was followed by a slight sea-level fall (∼10 m in the smoothed record) between ∼139 and 138 ka. Finally, a more significant pulse of ∼70 m in RSL rise (MWP-2B) is inferred between 135 and 130 ka. The period between the ephemeral pulse of sea-level rise at the beginning of the PDG (MWP-2A) and the second prolonged pulse (MWP-2B/HS11) has sometimes been referred to as the PDG sea-level reversal (Siddall et al.2006).

The coral RSL data from Huon Peninsula and Tahiti independently provide additional evidence for an ephemeral reversal in sea-level rise occurring during the penultimate deglaciation (Fig. 7). In the case of Tahiti, sedimentary evidence for the superposition of shallow and deeper water facies led to the interpretation that there was an ephemeral deepening (sea-level rise) followed by a return to shallower water conditions (sea-level fall or stabilization) (Thomas et al.2009). The Tahiti data provide bounding ages on the timing of this sea-level rise pulse, with ages of corals that grew at 135.0 ka (in 0–6 m water depth) and 133.5 (±1) ka (0–25 m water depth). In between these shallower facies, there is a deeper water facies (≥20 m paleo-water depth), but there are no reliable ages within this interval of the core (Thomas et al.2009). This observation, based on changes in both the lithofacies and benthic foraminiferal assemblage, is interpreted as a pulse of sea-level rise in between about 135.0 and 133.5 ka (Fujita et al.2010). A similar sea-level oscillation has also been interpreted based on the stratigraphy as well as the age and paleo-water depth reconstruction at Huon Peninsula (Esat et al.1999). The absolute timing of coral growth is only loosely constrained at this site due to the open-system behaviour of the U-series isotopes (as reflected by the scatter in ages of corals collected in Aladdin's Cave, ∼134 to 126 ka; Fig. 7) (Esat et al.1999). Indeed, the corals from Terrace VII have ages (with high uncertainty) ranging from about 137 to 134.5 ka and the corals from the cave have a wide range of ages, from 134.1 to 125.9 ka (more details in Esat et al.1999). Given that the younger end of this age range is clearly within the MIS 5e sea-level highstand (e.g. Stirling et al.1998), it is more likely that the older end of this diagenetic array of data from Aladdin's Cave is a better approximation for the primary age (i.e. it is closer to the unaltered endmember). Despite these diagenetic concerns, the agreement in the timing of this PDG sea-level reversal (MWP-2A) in Tahiti and Huon Peninsula is striking (Fig. 7b).

When considering the 95 % probabilistic intervals of the Red Sea RSL reconstruction on the chronology from Grant et al. (2012), an overlap is observed with the coral data over the MWP-2A interval, within the stated uncertainties. Still, both coral datasets suggest that MWP-2A occurs several millennia later (i.e. ∼135–134 ka) than in the Red Sea RSL reconstruction. This mismatch is likely to be related to the difficulty of precisely anchoring the dating of the current Red Sea RSL age scale over this interval (as also discussed in the Supplement of Grant et al.2012). Hence, we propose a revised chronology for the Red Sea RSL record in order to provide a better agreement with the absolutely dated corals. Given the potential ambiguities of the tie point defined in Grant et al. (2012) to stretch the depth scale across this interval, we find it reasonable to adjust it such that the timing of MWP-2A is more consistent with the absolute ages provided by the Tahiti and Huon Peninsula coral data.

We note that reassigning the tie points across this interval (Table S5), where tie points are placed at the beginning and end of MWP-2A (as defined by the coral data), results in a sea-level reconstruction that more closely approximates a linear age–depth model (Fig. 7a). This revised age model for the Red Sea RSL is adopted as our preferred reconstruction for sea-level change during the PDG. This reconstruction also compresses the total duration of the sea-level rise during the entirety of the PDG transition, which has implications for the freshwater forcing in the NH and for making analogies between the last and penultimate deglaciations.

This revised chronology is still attached to large uncertainties given the limits of the datasets. Also, considerable uncertainties remain with the magnitude of the sea-level pulse during MWP-2A because some of the corals cover a wide range of paleo-water depth (0 to 6 m for the pre-MWP-2A Tahiti corals, ≥20 m for the Tahiti corals during MWP-2A, 0–25 m for the post-MWP-2A Tahiti corals and 0 to 20 m for the Aladdin's Cave corals). Despite these uncertainties in the absolute position of sea level, the relative sea-level changes for each site clearly demonstrate an ephemeral deepening during meltwater pulse MWP-2A in both cases.

Glacial isostatic adjustment to the deterioration of the PGM ice sheets will also differentially affect Tahiti and Huon Peninsula, which precludes a direct comparison of the magnitude of sea-level change between these sites or a direct interpretation of global mean sea-level change in the absence of modelling. Because the changes in global mean sea level are rapid across the penultimate deglaciation, the eustatic signal is likely dominant, leading to a timing of the rapid changes that is similar between local RSL and global mean sea-level reconstructions. Still, the rate of change may be different between sites due to local differences in the magnitude of sea-level change. Based on the revised chronology for the Red Sea RSL and on the coral constraints, MWP-2A starts at ∼137 ka, while MWP-2B starts at ∼133 ka (Fig. 7, Table 4).

Finally, far-field coral data from the Seychelles and Western Australia, which have been corrected for the glacial isostatic adjustment (e.g. Dutton et al.2015), indicate that global mean sea level passed the position of modern sea level at about 129 ka (Fig. 7). The evolution of sea level during the LIG highstand is still debated and may have included some metre-scale sea-level oscillations, but by at least some accounts, it is thought to have risen a few metres between 129 and 122 ka (e.g. Kopp et al.2009; Dutton et al.2015). So while the timing of peak sea level may have occurred later in the interglacial (∼122 ka), the onset of the highstand (∼129 ka) could represent an inflexion point in the rate of sea-level change coming out of the rapid deglaciation and into the interglacial.

Overall, eustatic sea-level reconstructions based on paleo-data and continental ice-sheet simulations (Sect. 4) are consistent. However, the amplitude of the eustatic sea-level change across the PDG estimated from the Red Sea reconstructions is ∼10 m smaller than the combined ice-sheet simulations (Fig. 4e). In addition, the sea-level data suggest a small sea-level increase at ∼140 ka, which is not present in the ice-sheet simulations. Both suggest that the main phase of sea-level rise or continental ice-sheet disintegration initiates at ∼134 ka, even if the overall magnitude is larger in the ice-sheet simulation, but with a lower rate of change than in the Red Sea reconstructions. However, it is worth keeping in mind that both the sea-level data and ice-sheet-model-based approaches are associated with large uncertainties regarding the exact timing and amplitude of global sea-level changes across the PDG.

6 Protocol for transient simulations of the PDG
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To this date, no transient simulation covering the period 140 to 127 ka has been performed with a three-dimensional climate model. However, the climate evolution over the period 135 to 115 ka has been successfully simulated with the Earth system model LOVECLIM (Loutre et al.2014; Goelzer et al.2016a). Their simulations highlight the potential and feasibility of transient simulations of the PDG. In addition, as a proof of concept, transient experiments of the last deglaciation have been successfully performed with AOGCMs and Earth system models (Liu et al.2009; Menviel et al.2011; Roche et al.2011; Gregoire et al.2012; He et al.2013; Otto-Bliesner et al.2014). As detailed in the previous sections, the proposed transient simulation of the PDG will use boundary forcings consistent with the ones used for the last deglaciation (Ivanovic et al.2016): i.e. appropriate orbital parameters, greenhouse gases concentration, continental ice-sheet geometry and meltwater input (Fig. 1). To maximize the use of the transient simulations, the transient simulations of the penultimate and last deglaciations, as well as the piControl should be performed with the same version of the climate model.

6.1 Equilibrium spin-up at 140 ka (PDGv1-PGMspin)

If a LGM run is already available, then it is suggested to initialize the 140 ka spin-up from the climate fields and ocean state produced by the LGM equilibrium run. Starting from a LGM state may minimize the duration of the spin-up, as it should shorten the time to reach equilibrium in the deep ocean (Zhang et al.2013). If starting from a pre-industrial set-up, it is suggested to follow the PMIP4 LGM protocol (Kageyama et al.2017) to set up the 140 ka state, but using the 140 ka boundary conditions described below instead of the respective 21 ka boundary conditions.

The model should be forced with 140 ka background conditions (Table 1), including appropriate orbital parameters, GHG concentrations as averaged over the interval 141–139 ka (191 ppm CO2, 385 ppb CH4 and 201 ppb N2O), as well as the NH and Antarctic ice sheets' extent, topography, and associated albedo (as described above). The forcing file describing the evolution of NH and Antarctic ice sheets, including the evolution of the ice mask, as well as surface and bedrock elevations, as simulated by ice-sheet models (Abe-Ouchi et al.2013; Briggs et al.2014) (Sect. 4), is available on the Research Data Australia Repository. These sections include the evolution of the ice mask, as well as surface and bedrock elevations. Kageyama et al. (2017) provides guidelines for computing the land-ice fraction and orography from the ice-sheet reconstruction datasets. The details of these forcings and the approach taken to compute them will ultimately depend on each model resolution.

The large glacial ice sheets of the PGM impacted sea level and the land–sea mask. It would be best to modify the land–sea mask resulting from ice-sheet changes. Depending on the resolution of the model, this might not be a crucial parameter, except for some bathymetry and land–sea mask features, which have particular importance for ocean circulation. For example, the Bering Strait, which is 40 to 50 m deep, exerts a significant control on NADW and North Pacific Intermediate Water formation (e.g Okazaki et al.2010; Hu et al.2012). During the 140 ka spin-up, the Bering Strait should be closed. Following the recommendations for the PMIP4 LGM equilibrium run (Kageyama et al.2017), the land–sea mask should include the exposure of the Sahul and Sunda shelves in the Indo-Australian region, as well as closure of the strait between the Mediterranean Sea and the Black Sea.

To account for the maximum PGM ice-sheet expansions and associated global sea level of ∼100 m below pre-industrial times, global salinity should be set to +0.85 psu above pre-industrial level. Furthermore, if oxygen isotopes are included in the simulations, the ocean mean δ18O should be initialized at 1 ‰, and if a carbon cycle model is included, the global mean alkalinity content should be increased by about 80 µmol L−1. Compared to the LGM state, these values are 0.15 psu lower for global salinity, 0.2 ‰ lower for mean ocean δ18O and about 16 µmol L−1 lower for global alkalinity.

The model should be spun-up until near equilibrium is reached. Previous PMIP protocols recommend that the simulations are considered at equilibrium when the trend in globally averaged SST is less than 0.05 C per century and the Atlantic Meridional Overturning Circulation (AMOC) is stable. Marzocchi and Jansen (2017) recently pointed out that the AMOC should be monitored on a centennial timescale to properly assess equilibrium. Zhang et al. (2013) further suggested that the trend in zonal-mean salinity in the Southern Ocean (south of the winter sea-ice edge) should remain small (less than 0.005 psu per 100 years), especially in the Atlantic sector, to avoid potential transient characteristics in the deep ocean from impacting on AMOC strength. For models including representations of the carbon cycle or dynamic vegetation, the requirement is that the carbon uptake or release by the biosphere is less than 0.01 PgC per year. Similar to the recommendation made in Kageyama et al. (2017), the outputs of at least 100 years of the equilibrated 140 ka spin-up should be made available and fully described.

6.2 Transient forcings across the PDG (PDGv1)

The main changes in boundary conditions across the deglaciation, i.e. insolation, GHG concentrations and continental ice sheets, have been described in Sects. 2, 3 and 4, respectively and are summarized in Table 1. For all simulations, methods should be fully documented.

6.2.1 Orography, bathymetry, coastlines and rivers

Disintegration of continental ice sheets during the deglaciation affected continental topography and ocean bathymetry and thus coastal outlines and river routing. Therefore, time-varying changes in land-ice fraction, land–sea fraction, and topography should be applied. Variations in the ice mask and topography should be updated at the same time. It is up to each group to decide the appropriate time frequency at which to update this forcing. The resolution of the files provided is 500 years, but higher-frequency changes obtained through linear interpolations can also be performed to avoid step changes. As mentioned for the 140 ka spin-up regarding changes in land–sea fraction, particular attention should be given to the opening of the Bering Strait and the flooding of the Sunda and Sahul shelves. When possible, these should be varied across the PDG as ice sheets disintegrate and sea-level rises. Following the combined ice-sheet history presented here, which includes some glacial isostatic adjustment (GIA) adjustment, the Bering Strait might open at ∼127.5 ka. Please note that this is a first estimate, which is associated with large uncertainties.

As changes in the land–sea mask could impact water delivery to the ocean through rivers, River mouths should be consistent with the adjusted land–sea mask (Kageyama et al.2017). If possible and of interest, river networks could also be remapped to take into account the ice-sheet changes.

6.2.2 Vegetation, land surface and other forcings

The climatic and ice-sheet changes occurring between 140 and 127 ka will significantly impact the vegetation and thus also land albedo, evapotranspiration and the terrestrial carbon pool. The preferred option would thus be to include a dynamical vegetation model, fully coupled to the atmospheric model. Care should be taken in regions where an ice sheet is present, as the ice sheet and its albedo should replace any possible vegetation. If a coupled dynamical vegetation model is not available, then the experiments should be run with prescribed land-surface parameters and with fixed vegetation types and plant physiology outside of the regions covered by a continental ice sheet, as obtained from the CMIP5 pre-industrial set-up (Taylor et al.2012; Ivanovic et al.2016). In that case, care should be taken to have a consistent land-surface or vegetation forcing with the adjusted land–sea mask. The vegetation or land-surface type on newly emerged land (for example the Sunda shelf) will thus need to be fixed based on interpolations with the nearest grid points.

6.2.3 Freshwater forcing

Through their impact on global salinity and ocean circulation, disintegrating ice sheets can significantly affect the climatic and biogeochemical evolution of the penultimate deglaciation (Oppo et al.1997; Cheng et al.2009; Hodell et al.2009; Landais et al.2013; Deaney et al.2017). In particular, meltwater input in the North Atlantic region could be a significant driver of changes in NADW formation (Loutre et al.2014; Goelzer et al.2016b; Stone et al.2016), including the ones associated with HS11 (further discussed in Sect. 8.2). It is thus strongly recommended to include a carefully designed freshwater scenario when performing transient simulations of the deglaciation.

As much as possible, and for all scenarios, meltwater should be added in the appropriate locations to match the evolution of the ice sheets. Freshwater can be added over an appropriate ocean area close to the disintegrating ice sheet, or a self-consistent paleo-surface drainage forcing could also be implemented. This would involve using the provided downslope routing fields to route the water flux (fwf, cm yr−1) from each grid cell:


with P for precipitation (cm yr−1), E for evaporation (cm yr−1) and (dH∕dt)icesheet (cm yr−1, assuming an ice density of 0.91 g cm−3) for the change in ice-sheet thickness over time as described in the ice-sheet forcing files. Modellers may need to adjust the above to account for land-surface model changes in water storage. In the case of negative meltwater forcing, the artificial salt flux addition should be spread globally over the ocean.

Estimates of sea-level changes across the PDG suggest a global sea-level rise of ∼100 m (Grant et al.2012, 2014) (Fig. 7) between 140 and 130 ka, due to the disintegration of NH and Antarctic ice sheets. As the Antarctic contribution to sea level is estimated at ∼13 m (Fig. 4), this leaves a ∼87 m contribution from NH ice sheets.

For the NH, three meltwater scenarios are proposed based (i) on changes in NH ice sheets, as described in Sect. 4 (Fig. 4f, black, fIC) (Abe-Ouchi et al.2013), (ii) on global mean sea-level changes with our revised chronology (see Sect. 5, Fig. 4f, blue, fSL – meltwater forcing scenario based on sea-level changes) minus a linear Antarctic contribution of 13 m (Fig. 4d) (Briggs et al.2014), and (iii) as derived from North Atlantic and Norwegian Sea IRD records (see also Sect. 8.2, Fig. 4f, red, fIRD – meltwater forcing scenarios based on the IRD record).

Estimates of meltwater input to the North Atlantic based on changes in NH ice sheets (Fig. 4b, e, f, black, fIC) (Abe-Ouchi et al.2013) suggest a sustained (≥0.1 Sv) meltwater flux between ∼133 and 127 ka. This forcing closely follows changes in high northern latitude summer insolation and would probably lead to a significant NADW weakening during HS11, lasting until about 125.5 ka depending on the model's sensitivity.

Meltwater input estimates derived from Red Sea sea-level records (Fig. 4f, blue, fSL) on the revised chronology (Sect. 5) display a large (up to 0.3 Sv) 1000-year-long meltwater pulse centred at 137 ka, a broad meltwater input between 134 and 131 ka with a large peak at 131.7 ka, and two relatively late meltwater pulses centred at 130 and 128.3 ka, respectively. The magnitude and length of the AMOC perturbation resulting from such a meltwater scenario will, of course, depend on each model's sensitivity and on the initial AMOC state. However, for most models, it is expected that NADW formation would weaken significantly between ∼137.8 and 136.5 ka as well as between ∼133.5 and 129 ka. Another small AMOC perturbation is expected between ∼129 and 127.8 ka in this scenario.

There are significant uncertainties associated with both the simulation of NH ice sheets and the timing and amplitude of sea-level changes. In addition, to fully explore the potential of transient deglacial simulations, it is critical to simulate NADW changes in global agreement with proxy records. Since periods of increased IRD delivery have been associated with changes in NADW (e.g. van Kreveld et al.2000; Rodrigues et al.2017), we design an additional meltwater scenario, for which the timing is based on North Atlantic and Norwegian Sea IRD records (Fig. 8a). To construct the fIRD scenario, the IRD record of MD95-2010 (Risebrobakken et al.2007) is assumed to represent freshwater input only originating from the Eurasian ice sheet for the period 140 to 133.8 ka, while the stack of IRD records presented in Fig. 8b (black) is assumed to represent freshwater originating from both the North American and Eurasian ice sheets for the period 133.7 to 127 ka. The normalized MD95-2010 IRD record and the IRD stack were thus scaled so as to obtain a total NH sle contribution of 87 m (Fig. 4e, red), with 35 m originating from the Eurasian ice sheet and 51 m from the North American one. It is expected this scenario will lead to a weakening of NADW between 139.5 and 136.5 ka as well as at ∼135 ka. The sustained meltwater pulse might induce NADW cessation between ∼133.5 and 129.4 ka, followed by a recovery sometime between 129 and 128 ka.

Figure 7Sea-level changes across the PDG: (a) stacked LR04 benthic δ18O curve (turquoise) (Lisiecki and Raymo2005) and Red Sea relative sea-level (RSL) data (1 kyr moving Gaussian filter in dark blue line and 95 % probability interval in light blue) in the age model from Grant et al. (2012) and in the new age model (this study, data in grey, 1 kyr moving Gaussian filter in black); RSL curve inferred from Red Sea Geo-KL11 and MD92-1017 cores on a depth scale (dark grey) (Grant et al.2012). (b) Tahiti (orange dots) (Thomas et al.2009) and Huon Peninsula (green dots) (Esat et al.1999) corals superimposed onto the Red Sea RSL curve displayed in the new age model (black). Orange bars indicate the range of the paleo-water depth estimates from Tahiti corals. Coral data have been updated to new decay constants (Cheng et al.2013) (Tables S3, S4). Black dots indicate the tie points defined so that the timing of MWP-2A in the Red Sea RSL is consistent with the absolutely dated Tahiti and Huon Peninsula coral data (Table S5). The pink box represents the timing and range of RSL during the main phase of LIG reef growth (129–122 ka) (Dutton and Lambeck2012). Vertical dashed grey lines indicated the 140, 135 and 130 kyr time intervals in both panels.


Figure 8Selection of RSL and North Atlantic marine sediment records: (a) Red Sea RSL record (Grant et al.2012), with revised age scale as described in Sect. 5. (b) Normalized IRD stack (black) of cores CH69-K09 (Labeyrie et al.1999), ODP980 (Oppo et al.2006), ODP983 (Barker et al.2015), ODP984 (Mokeddem et al.2014), ODP1063 (Deaney et al.2017) and SU90-03 (Chapman and Shackleton1998) with ±1σ (grey envelope). Normalized IRD record of MD95-2010 (blue) (Risebrobakken et al.2006). (c) Neogloboquadrina pachyderma sin. (%) in cores ODP980 (cyan) (Oppo et al.2006), ODP983 (green) (Barker et al.2015) and ODP984 (magenta) (Mokeddem and McManus2016). (d) Estimated summer SST anomalies (C) in cores ODP980 (cyan), CH69-K09 (black) (Labeyrie et al.1999) and SU90-03 (blue) (Chapman and Shackleton1998). (e) U37k SST anomalies (C) in cores MD01-2444 (red) (Martrat et al.2007), ODP976 (magenta) and MD95-2042 (grey) (Martrat et al.2014); (f) percentage of warm (circles) and cold (axis reversed) species in core ODP1063 (Deaney et al.2017). (g) Estimated δ18Osw () in cores ODP976 (red), MD95-2042 (magenta) (Martrat et al.2014), SU90-03 (blue) (Chapman and Shackleton1998), CH69-K09 (black) (Labeyrie et al.1999), ODP984 (green) (Mokeddem and McManus2016) and ODP980 (cyan) (Oppo et al.2006); Benthic δ13C () in cores (h) ODP983 (green) (Lisiecki and Raymo2005) and ODP980 (cyan) (Oppo et al.2006). (i) MD95-2042 (black) (Martrat et al.2014). (j) a δ13C stack of cores U1308, CH69-K09 and ODP1063 (Labeyrie et al.1999; Hodell et al.2008; Deaney et al.2017). All the age models have been revised as described in the Methods section and in Govin et al. (2015).


As can be seen in Fig. 4f, fSL and fIRD share some similarities. The main differences between these scenarios are the small meltwater pulse at ∼137 ka in fSL, which is of much reduced amplitude in fIRD, and the ∼128 ka pulse in fSL not present in fIRD. Finally, the fSL scenario includes periods of significant negative meltwater forcing (i.e. artificial salt flux addition), corresponding to phases of sea-level lowering. As described above, it is suggested to use the self-consistent drainage scheme for the location of the meltwater input. For NH scenarios fSL and fIRD, (dH∕dt)icesheet (from Eq. 1) would need to be scaled to obtain the appropriate meltwater flux.

For those who wish to take part in an inter-comparison involving comparable boundary conditions, the fSL scenario is put forward as the recommended option (Table 1). Depending on the sensitivity of the model to freshwater input, the fIRD scenario can provide a good alternative to fSL. However, the ultimate choice of the appropriate freshwater scenario is left to each group, and sensitivity experiments to assess the climatic impact of different meltwater scenarios are encouraged (Sect. 7).

To take into account the effect of Antarctic ice-sheet melting, freshwater should also be added close to the Antarctic coast, following the self-consistent routing scheme described above (Fig. 4d) (Briggs et al.2014). This scenario will broadly consist in adding ∼0.0135 Sv meltwater from 140 to 130 ka. However, there are significant uncertainties associated with the timing of the Antarctic deglaciation. Additional experiments are necessary to further constrain the impact of the Antarctic ice-sheet disintegration on the deglacial climate and carbon cycle (e.g. Menviel et al.2010). Therefore, another scenario inspired by the Antarctic ice-sheet deglacial trajectory described in Goelzer et al. (2016a) is proposed (Table 2, fSL2).

Finally, if preferred and instead of the Northern and Southern Hemisphere freshwater scenarios described here, a globally uniform freshwater flux that corresponds to the ice-sheet evolution can be added to simply conserve salinity throughout the transient deglacial experiment (Table 2, fUN). This latter option is equivalent to the melt-uniform scenario used in the PMIP4 transient simulations of the last deglaciation (Ivanovic et al.2016).

Table 2Freshwater scenarios for transient simulations of the PDG. Freshwater can be added in the NH and close to the Antarctic coast based on meltwater routing or as a globally uniform flux. The Antarctic freshwater forcing in fSL2 is based on Goelzer et al. (2016a) but scaled down to obtain a total Antarctic contribution of 20 m sle.

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6.3 Requested variables

A transient simulation of the PDG implies that the climate model will need to be integrated for 13 000 years, thus generating a large amount of data. Depending on the spatial resolution of the different model components, storage of data outputs can become an issue. We therefore suggest a list of variables that absolutely needs to be uploaded to the Earth System Grid Federation (ESGF) database (Balaji et al.2018) to understand the results and perform inter-model and model–data comparisons. The CMIP6 conventions should be used for all the variable names and units. Table 3 presents a list of forcings, and variables related to the atmosphere, ocean, vegetation and ocean biogeochemistry realms (if relevant) to be saved either as annual or monthly means. In addition, to this list standard global, hemispheric or zonally averaged variables would need to be included, such as northern and southern sea-ice extent and surface air temperature, maximum meridional overturning stream function in the North Atlantic, top of the atmosphere (TOA) energy budget, surface energy budget, carbon budget (if relevant), etc... The list detailed in Table 3 represents the strict minimum of variables to be distributed. It is expected that additional variables will be included by each group, particularly if other model components are included (e.g. isotopes).

Table 3Requested one-dimensional(1), two-dimensional (longitude, latitude(X,Y); latitude, basin(Y,B); or latitude, depth (Y,Z)) and three-dimensional (latitude, longitude, depth or basin, longitude, depth; unmarked in the table) variables to be uploaded to the ESGF database. Most variables are requested as annual means, but * indicates that monthly outputs are requested. The CMIP6 conventions should be used for the variable names and units. See (last access: May 2019) for details.

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7 Sensitivity experiments
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The penultimate deglaciation is a particularly interesting period as it provides a framework to study the impact of changes in insolation, atmospheric GHG concentrations and continental ice sheets on climate. In addition, meltwater input associated with the disintegration of continental ice sheets will also impact the oceanic circulation and thus global climate and biogeochemistry (e.g. Liu et al.2009; Menviel et al.2011, 2014; Schmittner and Lund2015; Goelzer et al.2016a; Ivanovic et al.2017; Menviel et al.2018). Ultimately, transient deglacial simulations will provide information on the impact of each of these processes as well as their interactions.

However, the transient simulation of the PDG proposed here might present a challenge to state-of-the-art Earth-system models, as they will need to include a spin-up with 140 ka boundary conditions and be run for 13 000 years. For this reason (i.e. to avoid the necessity for additional simulations when the computational expense is prohibitive), the main experiment includes all appropriate boundary forcing as well as meltwater input due to disintegrating ice sheets. This experiment will provide valuable information on processes occurring during the PDG and will allow for a thorough comparison of the penultimate and the last deglaciation by complementing existing transient simulations of the last deglaciation (Liu et al.2009; Menviel et al.2011; Roche et al.2011; Gregoire et al.2012; He et al.2013) and the new experiments performed as part of PMIP4 (Ivanovic et al.2016). These transient simulations of the PDG will also complement existing transient simulations of the PDG performed with the LOVECLIM Earth-system model and covering the time interval 135 to 115 ka (Loutre et al.2014; Goelzer et al.2016a). Finally, the proposed experiment will provide a link to the PMIP4 transient simulation of the LIG (127 to 121 ka) as well as the PMIP4 127 ka time slice experiment (lig127k) (Otto-Bliesner et al.2017), even though the protocol of the LIG experiments includes pre-industrial continental ice sheets.

The main experiment is described in Table 1 and includes a comprehensive meltwater scenario. Additional transient simulations of the PDG are encouraged, for example, to assess different timing and amplitude of meltwater input (Table 2) but also simulations with globally uniform meltwater input (fUN). As there are large uncertainties associated with meltwater input scenarios and the sensitivity of deep convection to the freshwater input, it is strongly advised to perform both the main experiment and fUN to isolate the impact of the freshwater input (Goelzer et al.2016a). In addition, the response to individual forcings (i.e. orbital parameters, GHGs and ice-sheet extent and albedo) could be assessed separately (He et al.2013; Gregoire et al.2015).

Even though there are some geological constraints on glacial evolution (see Sect. 4.1.), there remain large uncertainties associated with the reconstruction of continental ice sheets during the PGM, across the PDG and during the LIG. In addition, there are significant uncertainties associated with the parametrization of dynamical processes governing continental ice sheets and most importantly those representing the climate forcing. Through their impact on albedo and topography, continental ice sheets can significantly influence climate (e.g. Timm et al.2010; Zhang et al.2014), even when the ice sheets are small (e.g. Löfverström et al.2014; Gong et al.2015; Roberts and Valdes2017; Gregoire et al.2018). Therefore, the impact of ice-sheet extent and topography on the PGM, across the PDG and during the LIG should be studied in detail through sensitivity experiments.

Models that include prognostic aerosol and/or dust could provide very useful fields to the community. Similarly, evaluating the impact of varying deglacial dust levels on climate and biogeochemistry are of crucial importance (Evan et al.2009; Martínez-García et al.2014). Therefore, sensitivity simulations forced with different dust-flux scenarios are encouraged.

8 Comparing model simulations to paleo-climate and paleo-environmental reconstructions
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It is central to PMIP to evaluate the realism of the coordinated transient simulations and the associated sensitivity experiments with environmental and climate reconstructions from archives. In the following, we first report on the few existing surface temperature syntheses that cover at least part of our interval of interest. We then provide a non-exhaustive selection of additional paleo-climatic and paleo-environmental records extending back to 140 ka. This will allow a first-order comparison between the changes recorded in different parts of the climate system and those inferred in the transient simulations (Figs. 8 and 9, Tables 4 and 5). The selection criteria are related to each record's temporal resolution and to the climatic or environmental interpretation that can be inferred from the measured tracer. We put a special emphasis on selecting marine sediment core records that can provide information on millennial-scale changes occurring in the North Atlantic sector during HS11, potentially linked with changes in NADW formation. This section ends with a review of the main limitations that will be associated with the model–data comparison and recommendations.

Grant et al. (2012)Raymo et al. (2004)Barker et al. (2015)Oppo et al. (2006)Shackleton et al. (2003)Hodell et al. (2008)Labeyrie et al. (1999)Deaney et al. (2017)Govin et al. (2009)Govin et al. (2012)Oppo et al. (2006)Shackleton et al. (2003)Martrat et al. (2014)Drysdale et al. (2009)Tzedakis et al. (2018)Marino et al. (2015)

Table 4Key paleo-proxy records of changes in ocean oxygen (δ18O) and carbon (δ13C) isotopic composition. The chronology of all paleo-proxy records presented here is based on an alignment with a Corchia U-Th-based chronology (Table S1). Letters in brackets indicate the major changes identified in the paleo-climatic records shown in Fig. 9. The dates of the major changes obtained through RAMPFIT or BREAKFIT are indicated with their associated uncertainties. Age uncertainties (1σ) reported include (1) the “internal” error of the event given by RAMPFIT (for most major changes) (Mudelsee2000) or BREAKFIT (major change dates highlighted with **) (Mudelsee2009), (2) the relative error related to the climatic alignment method for marine sediment records, and (3) the absolute dating error of the reference timescale. Based on North Atlantic records, changes have been split into five phases (φ), with the date representing the beginning of each phase: φ1 is associated with the early phase of HS11 and MWP-2A, φ2 represents a pause within HS11, φ3 the main phase of HS11 and MWP-2B, φ4 the inception out of HS11, and φ5 full interglacial conditions. Implicitly, the end of each phase corresponds to the beginning of the next one. For the sea-level record, the new chronology discussed in Sect. 5 is used. CS98 refers to Chapman and Shackleton (1998). “Ventil.” stands for “ventilation”.

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Bereiter et al. (2015)Oppo et al. (2006)Cortijo et al. (1999)Martrat et al. (2014)Govin et al. (2009)Govin et al. (2012)Pahnke et al. (2003)Jouzel et al. (2007)Drysdale et al. (2019)Brauer et al. (2007)Tzedakis et al. (2003)Cheng et al. (2016)Yang and Ding (2014)

Table 5Same as Table 4 for key paleo-proxy records of climatic and environmental changes through the PDG selected for their relatively high resolution. An indication of their climatic or environmental use is indicated in italics and the proxy used is shown in parenthesis in column 1. Age uncertainties (1σ) reported include (1) the “internal” error of the event given by RAMPFIT (for most major changes) (Mudelsee2000) or BREAKFIT (major change dates highlighted with **) (Mudelsee2009), (2) the relative error related to the climatic alignment method for marine sediment and Ioannina records, and (3) the absolute dating error of the reference timescale. No dating error is provided in Yang and Ding (2014). As a result, the stated error here only encompasses the internal error; it thus represents only a minimal error for the timing of the stacked grain size increase. The mean age representing the beginning of each phase (φ) is shown in the last row and is calculated from all the available dates within each phase shown in Tables 4 and 5. FFA stands for foraminifera faunal assemblage. CS98 refers to Chapman and Shackleton (1998). Based on North Atlantic records, changes have been split into five phases (φ), with the date representing the beginning of each phase: φ1 is associated with the early phase of HS11 and MWP-2A, φ2 represents a pause within HS11, φ3 is the main phase of HS11 and MWP-2B, φ4 is the inception out of HS11, and φ5 is full interglacial conditions. Implicitly, the end of each phase corresponds to the beginning of the next one.

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Figure 9Selection of paleo-records illustrating climatic and environmental changes between 140 and 127 ka. (a) SST records from North Atlantic cores ODP-976 (pink) (Martrat et al.2014) and SU90-03 (blue) (Chapman et al.2000). (b) Percentage of temperature tree pollen from the Greek Ioannina sequence (dark green) (Tzedakis et al.2003) and from the Italian Lago Grande di Monticchio (LGdM) sequence (light green) (Brauer et al.2007). (c) Composite speleothem calcite δ18O record from the Italian Corchia Cave speleothem (purple) and Corchia Cave deposit Mg concentration Mg concentration (Drysdale et al.2007, 2019) (d) Composite calcite δ18O record from the Chinese Hulu, Sanbao and Dongge caves speleothems (orange) (Cheng et al.2016). (e) Stacked grain size record of Chinese loess (light grey) (Yang and Ding2014). (f) SST records from Southern Ocean cores MD97-2120 (light blue) (Pahnke et al.2003) and MD02-2488 (dark blue) (Govin et al.2012). (g) δ13C record from North Atlantic core MD95-2042 (light pink) (Shackleton et al.2003) and Southern Ocean core MD02-2488 (dark blue) (Govin et al.2012). (h) Antarctic ice δD record from EPICA Dome C (Jouzel et al.2007) on the AICC2012 chronology (black) (Bazin et al.2013; Veres et al.2013). Letters and yellow and black crosses indicate the major changes identified in the paleo-climatic records and their ±1σ dating uncertainty (horizontal grey bars, Tables 4 and 5). Chronologies and associated references are detailed in Tables 4 and 5. No details on the dating errors attached to the Chinese loess stacked grain size record are provided in Yang and Ding (2014); the 1σ error should thus be treated as an underestimated dating error.


8.1 Available surface temperature syntheses

Quantitative comparisons between paleo-reconstructions and model outputs across the time interval 140–127 ka are possible but remain limited to a few parameters (e.g. surface air and sea surface temperature, SAT and SST). Qualitative and indirect comparisons are also adequate to evaluate simulations: for example, simulated AMOC strength against paleo-records (Sect. 8.2), simulated precipitation compared to Chinese speleothem calcite δ18O records or the simulated vegetation patterns against pollen-based vegetation reconstructions (Sect. 8.3, Table 5).

There are currently no paleo-climate data compilations focusing on the PGM or covering the full length of the PDG, but syntheses of quantitative and temporal surface temperature changes focusing on the LIG are available for model evaluation of the second part of the penultimate deglaciation. Indeed, the synthesis by Capron et al. (2014) covers the time interval 135–110 ka and, in a coherent temporal framework underpinned by the AICC2012 ice-core chronology, compiles annual SAT above Greenland and Antarctica and summer SST records from the North Atlantic and the Southern Ocean located at latitudes above 40. In Uemura et al. (2018), updated surface air temperature reconstructions displayed on AICC2012 are presented for the EDC, Dome F and Vostok deep ice cores. They represent useful constraints, particularly to investigate spatial differences in temperature between East Antarctica inland ice-core sites during the PDG.

Further, based on harmonized chronologies between marine records, Hoffman et al. (2017) propose a global annual SST compilation with time series encompassing 130 to 115 ka. Hoffman et al. (2017) use the SpeleoAge age scale as a reference chronology. This SpeleoAge age scale results from the adjustment of the EDC3 ice-core timescale using radiometric dates from Chinese speleothems (Barker et al.2011) and presents age differences across the PDG with AICC2012 (e.g. ∼1.4 at 127 ka, ∼1.0 at 136 ka). Both syntheses are useful to perform site-to-site model–data comparisons in order to provide detailed information about the spatial structure of the changes. The regional stacks produced by Hoffman et al. (2017) provide a first-order estimate of the mean annual SST responses between 130 and 127 ka.

8.2 North Atlantic records to provide information on Heinrich stadial 11 and NADW changes

Deglaciation of NH ice sheets led to increased meltwater input into the North Atlantic, thus reducing surface water density and potentially weakening NADW formation. It has been shown that changes in NADW have a significant impact on North Atlantic and European climate (e.g. Stouffer et al.2007; Martrat et al.2014; Wilson et al.2015) but also have a strong imprint on tropical hydrology (e.g. Arz et al.1998; Chiang and Bitz2005; Deplazes et al.2013; Otto-Bliesner et al.2014; Jacobel et al.2016). Through oceanic and atmospheric teleconnections, NADW variations can further modulate the strength of the Asian Monsoon (e.g. Cheng et al.2009) and the formation of other oceanic water masses, such as North Pacific Intermediate Water (Okazaki et al.2010). Using a multi-proxy approach, we investigate here the millennial-scale variability occurring during the PDG and particularly potential changes in NADW linked with the meltwater input of HS11.

We selected 11 marine sediment cores from the western Mediterranean Sea, the North Atlantic and the Nordic Seas with relatively high-resolution proxy data across the PDG (Tables 4 and 5, Fig. 8). These sites have undergone multi-proxy investigations (e.g. stable isotopes on benthic and planktonic foraminifera, foraminifera faunal assemblages, IRD counts, SST reconstructions, Pa∕Th, ϵNd). The chronology of all the sites has been revised here using the radiometrically dated timescale of the Italian Corchia cave (original record in Drysdale et al.2009, see also Tzedakis et al.2018, for the high-resolution calcite δ18O profile) as a reference for the PDG (Sects. S1 and S2, Table S1 in the Supplement).

Phases of meltwater input and/or changes in NADW can be investigated using different proxies. The intensity of NADW formation can be deduced from variations in Pa∕Th (a kinematic proxy for the rate of deep water export from the North Atlantic) and ϵNd (a water provenance tracer) in North Atlantic sediment cores (McManus et al.2004; Roberts et al.2010), but only one record (ODP 1063) currently covers the PDG, and the time resolution of Pa∕Th and ϵNd measurements across the PDG is relatively low (Böhm et al.2015; Deaney et al.2017). Phases of iceberg discharges, which might significantly impact NADW formation, can be inferred from the amount of IRD in North Atlantic cores. For this time period, IRD records with sufficient resolution are available from six Atlantic cores (CH69-K09, ODP980, ODP983, ODP984, SU90-03 and ODP1063) and one Norwegian Sea core (MD95-2010). The IRD records were first normalized, after which an IRD stack was made for the North Atlantic cores by interpolating them onto a common age scale with a regular (100-year) time step.

Because continental ice sheets are 18O-depleted, the meltwater supply also induces a drop in seawater δ18O (δ18Osw), particularly at the ocean's surface. For six of the sites, changes in δ18Osw have been reconstructed from paired SST and planktic δ18Oc records using the following paleo-temperature equation (Shackleton1974):


with Tiso the isotopic or calcification temperature (C) that is deduced from SST reconstructions, δ18Oc the isotopic composition of the calcite (, PDB), and δ18Osw the isotopic composition of seawater (, standard mean ocean water; SMOW). The factor 0.27 is added for calibration against international standards (SMOW vs. Pee Dee Belemnite – PDB) (Hut1987).

As a weakened NADW transport leads to the accumulation of remineralized carbon below 2500 m in the North Atlantic (Marchal et al.1998), changes in oceanic circulation are also inferred from benthic foraminifera δ13C, a tracer for the ventilation of deep-water masses (Duplessy et al.1988; Curry et al.1988; Menviel et al.2015; Schmittner et al.2017). Finally, since an NADW weakening reduces the meridional heat transport to the North Atlantic (Stouffer et al.2007), it is generally accompanied by a sea surface cooling across this region (Kageyama et al.2013; Ritz et al.2013).

The Norwegian Sea core MD95-2010 (Risebrobakken et al.2006) indicates an increased IRD content starting at 139.5 ka (Fig. 8b). This could suggest an initiation of the penultimate deglaciation by enhanced iceberg calving (and subsequent melting) from the Fennoscandian ice sheet. However, this has little to no effect in the North Atlantic. Only in the western side of the North Atlantic, right in the meanders of the Gulf Stream, does core CH69-K09 display a surface cooling and a drop in δ18Osw starting at ∼137.5 ka (Labeyrie et al.1999).

A more significant deglacial pulse, indicated by an IRD increase in both the Nordic Seas and the North Atlantic, as well as a surface freshening in all North Atlantic cores occurs at ∼136.4–133.9 ka (Fig. 8g). This meltwater input, which could correspond to the first phase of HS11, could have led to a significant weakening of NADW, as indicated by ϵNd and Pa∕Th records from the Bermuda rise (Böhm et al.2015) (not shown), as well as by the accompanying surface cooling in the North Atlantic (Chapman and Shackleton1998; Martrat et al.2007, 2014; Deaney et al.2017) and a decrease in benthic δ13C below 2000 m depth (Labeyrie et al.1999; Shackleton et al.2003; Hodell et al.2008; Deaney et al.2017). The low-resolution ϵNd and Pa∕Th records from the Bermuda rise (Böhm et al.2015) also indicate a weakening of NADW transport during this period (not shown). This deglacial phase identified in North Atlantic marine sediment cores is broadly coincident with MWP-2A (Fig. 7, Sect. 5).

Martrat et al. (2014) identified a double-u structure in SST records from the western Mediterranean Sea (ODP976) and the Portuguese margin (MD95-2042, MD01-2444) during the PDG, with a short-duration warming centred at ∼133.6 ka. This warming is also seen in other North Atlantic records, such as ODP 984, 983, 980 and 1063 (Fig. 8c–f). Very interestingly, and within dating uncertainties, this event corresponds to a small increase in atmospheric CH4 (Figs. 1 and 2) (Landais et al.2013) and broadly corresponds to a pause in the sea-level rise. In parallel, Pa∕Th in core ODP1063 displays a significant decrease at this time (not shown), and benthic δ13C increases in the deepest North Atlantic cores (Fig. 8). This warming could thus be due to a short-lived reinvigoration of NADW formation.

In agreement with previous studies (e.g. Marino et al.2015; Tzedakis et al.2018), the main phase of HS11 occurs between ∼133.3 and 130.4 ka and is characterized by a large IRD peak, cold surface water conditions, a minimum in seawater δ18O, and low benthic δ13C values at most sites of the North Atlantic (Fig. 8). Elevated rates of iceberg calving and melting of NH ice sheets thus lead to a large meltwater pulse in the North Atlantic, a significant global seal level rise (MWP2-B) (Grant et al.2012, 2014), NADW weakening (Böhm et al.2015) and a surface cooling that is at least regional.

The resumption of NADW formation at the end of HS11 marks the end of the penultimate deglaciation (Tzedakis et al.2012) (∼130.4–128.5 ka). It is characterized by an increase in SST, seawater δ18O (Oppo et al.2006; Mokeddem and McManus2016; Martrat et al.2007, 2014; Deaney et al.2017) and benthic δ13C from the North Atlantic region (Fig. 8). The associated atmospheric warming in the North Atlantic region also leads to terrestrial regrowth and thus a sharp increase in atmospheric CH4 occurring between ∼129 and 128.5 ka (Landais et al.2013).

8.3 Other environmental and climate reconstructions

Figure 9 shows a non-exhaustive selection of other environmental and climate proxy records. Tables 4 and 5 further provide the climate or environmental interpretation for each selected tracer to aid the model–data comparison, including the tracers described in Sect. 8.2. Only a few speleothem calcite δ18O (δ18Oc) records and their environmental or climatic interpretation are presented here. However, it can be complemented with the compilation of speleothem δ18Oc records covering the time interval 140 to 110 ka presented by Govin et al. (2015). It is also important to recognize that the dominant drivers of speleothem δ18Oc may change over time and differ from one cave site to another (see Sect. 8.4). Therefore we strongly advise a critical assessment of these interpretations based on the most recent developments and advances in stable-isotope hydrology and the original publications.

Tables 4 and 5 also detail the timing of the major changes (and associated uncertainties) as recorded in each time series. These major changes are identified using the RAMPFIT or BREAKFIT software programs (Mudelsee2000, 2009). Age uncertainties (1σ) reported in Tables 4 and 5 include (i) the “internal” error of the event given by RAMPFIT or BREAKFIT, (ii) the relative error related to the climatic alignment method for marine sediment and Ioannina records, and (iii) the absolute dating error of the reference timescale. The selected ice- and sediment-based records are all displayed on AICC2012 or on a timescale coherent with this ice-core age scale, while terrestrial records are displayed on their own independent chronology. Among these paleo-climatic time series, several isotopic records are presented here and we strongly encourage transient simulations to be performed with oxygen and/or carbon-isotope-enabled models to allow direct quantitative comparison between simulated and measured isotopic time series.

Little change occurs until the beginning of phase 1 at ∼136.4 ka, after which a cooling phase is identified in a few records of the North Atlantic (Figs. 8 and 9a, major change (A)). This also corresponds to reduced monsoon activity as recorded in Chinese speleothems (Fig. 9d (M)), and the initiation of the Antarctic warming (Fig. 9h (W)). The short-lived warming event in the North Atlantic associated with phase 2 (Fig. 9a (B)) is also identified in the Chinese speleothems as a slightly wetter interval (Fig. 9d (N)). Other environmental records might not have the necessary resolution to record this multi-centennial-scale event. The main phase of HS11, corresponding to phase 3, is associated with meltwater input and cold conditions in the North Atlantic (Fig. 9a (C) and 9c (I)), dry conditions over Europe (Fig. 9b) and Asia (Fig. 9d, interval between (O) and (P)), and warmer conditions at high southern latitudes (Fig. 9f, h). The end of HS11 (phase 4) associated with a pause in the meltwater input (Fig. 9c (J)) and progressively warmer conditions in the North Atlantic and southern Europe (Fig. 9a (D, F) and 9c (K)) corresponds to a strengthening of the Asian monsoon (Fig. 9d (P) and 9e (Q)), and maximum warmth at high southern latitudes (Fig. 9f (R, S) and 9h (X)). Interglacial conditions in atmospheric CO2 and CH4 as well as North Atlantic temperatures and ventilation are attained at about 128.5 ka (Fig. 8), which is also associated with warm and wet conditions in southern Europe (Fig. 9b (G) and 9c (L)).

8.4 Limitations and recommendations

One important consideration to account for when comparing simulated variables against paleo-data between 140 and 127 ka is the large uncertainties associated with both absolute and relative chronologies of most paleo-climatic records during this time interval. Dating uncertainties range from a few centuries to up to several thousand years depending on the type of archive and dating methods (Govin et al.2015). For instance, the average absolute dating error attached to the Corchia Cave δ18O record is ∼0.7 ka (2σ) (Tzedakis et al.2018). For marine sediment chronologies, which are mainly based on record alignment strategies (e.g. a record on a depth scale is aligned onto a dated reference record), the age uncertainties encompass a relative dating error (“alignment error”) in addition to the absolute error attached to the chronology of the dated reference record. As a result, the overall 2σ age error associated with North Atlantic sediment core ODP976 aligned onto the Corchia record is 1.6 kyr on average (Table S2). Between 140 and 127 ka, the average absolute error attached to the AICC2012 chronology used to display ice and gas records from the EDC ice core is about 4 ka (2σ) (Bazin et al.2013; Veres et al.2013). This large AICC2012 dating uncertainty is thus attached to the GHG concentration records used to force the transient simulations (Sect. 3). It will therefore taint the relative timing of the changes in orbital and atmospheric CO2 forcing that will largely drive the simulated evolution of climate and environmental changes across the PDG. The relative timing of those changes will also be affected by uncertainties attached to the temporal evolution of the ice sheets and related meltwater forcing scenarios. However, uncertainties attached to the relative timing of changes between GHG forcing and the simulated changes in Antarctica and the Atlantic Ocean basin are somewhat reduced since the ice- and sediment-based records are also displayed on the AICC2012 timescale or on timescales coherent with the reference ice-core age scale (Sect. 8.2).

Limitations are also attached to the potential misinterpretation of climate and environmental proxies due to an incomplete understanding of how some of those archives record climatic and environmental change. First, SST records highlighted here have been reconstructed using various microfossil and geochemical methods. Although the use of various tracers is known to yield SST discrepancies in particular above 35 N (MARGO project members2009), the extent to which these different SST reconstruction methods influence the representation of temporal climatic changes across our studied time interval is poorly known. Additional difficulties arise from the individual methods commonly used to reconstruct past SST due to, for example, the poor understanding of the modern habitat (e.g. living season and water depth) of microfossil species (e.g. foraminifera) (Jonkers and Kučera2015) and alkenone producers (e.g. Rosell-Melé and Prahl2013). With these limitations acknowledged, model–data comparisons should use annual or appropriate seasonal climatic variables depending on the interpretation of the measured climate proxy proposed in the original or subsequent publications.

Regarding SAT reconstructions over Antarctica, the impact of changes in seasonality of snow precipitation on the reconstruction remains difficult to quantify based on ice-core water-isotopic records (e.g. Masson-Delmotte et al.2011; Uemura et al.2012). In addition, Sime et al. (2009) suggested that the temperature at Dome C should be much higher than the one inferred from water isotopes and a constant temperature–δ18O slope during MIS 5e. For now, we suggest the use of simulated annual mean climatic variables for the comparison. However, it is crucial that the seasonality of paleo-records is better assessed to improve the interpretation of temperature reconstructions and hence the model–data comparisons.

In addition, uncertainties remain regarding the dominant controlling factors (i.e. changes in temperature, rainfall amount and rain sources) of speleothem calcite isotopic records (e.g. Govin et al.2015). For instance, δ18Oc records throughout Asia are commonly interpreted as tracers of past changes in the intensity of the Asian monsoon. In particular, Chinese speleothem δ18Oc is classically interpreted as reflecting the East Asia monsoon (e.g. Cheng et al.2009, 2016). However, a water-hosing experiment performed with an oxygen-isotope-enabled model suggests instead that δ18Oc variations may reflect changes in the intensity of the Indian, rather than East Asian, monsoon precipitation during Heinrich events (Pausata et al.2011; Caley et al.2014). Another recent model study also demonstrates that variations in ice core and speleothem oxygen isotope reconstructions cannot solely be attributed to climatic effects but also reflect depleted δ18O of nearby oceans during glacial meltwater events (Zhu et al.2017). Overall, we strongly encourage the use of isotope-enabled models to allow for direct and quantitative model–data comparison of isotopic tracers.

Additional multi-centennial-scale paleo-climatic reconstructions are necessary in order to further constrain the millennial-scale variability during the PDG. It is also crucial that comprehensive data compilation work is carried out over the entire studied time interval and covering, in particular, the PGM in order to test the robustness of the initial 140 ka spin-up climate. Modelling groups running transient deglacial simulations and/or associated sensitivity experiments are encouraged to use multiple paleo-records for a full diagnosis of the simulations.

9 Conclusions
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Here, we present a protocol for performing transient simulations of the PDG spanning 140 to 127 ka. Changes in boundary conditions across the PDG that will serve as a forcing are presented and discussed. This includes changes in orbital parameters, GHG concentration, NH and Antarctic ice sheets, and associated deglacial meltwater input. While not used as a direct forcing, changes in global sea level are also presented on a new chronology. Finally, a series of key paleo-climatic and paleo-environmental records are suggested to perform model–data comparisons. The marine records were recovered from the North Atlantic and Southern Ocean, while the continental records were retrieved from Europe, China and Antarctica. Performing transient simulations with oxygen- and/or carbon-isotope-enabled Earth-system models could significantly improve model–data comparisons by providing a more direct and quantitative comparison with paleo-proxies based on measured isotopic signatures (e.g. δ18Oδ13C).

Simulations of the penultimate deglaciation would allow a comparison with the last deglaciation, therefore highlighting similarities and differences between the last two deglaciations. The evolution of insolation across the two deglaciations is different, potentially explaining the relatively more rapid disintegration of NH ice sheets during the PDG compared to the last deglaciation. Acting both as a response to and driver of changes, atmospheric CO2 appears to increase much more gradually during the penultimate than the last deglaciation. Another striking difference between the penultimate and the last deglaciation is the lack of a Bølling–Allerød warming in the middle of the deglaciation, even though, as discussed in Sects. 5 and 8, several records show a brief reversal centred at about 133.6 ka. Transient simulations can thus shed light onto the different and interactive roles of radiative forcing (insolation, GHG concentrations) and ocean circulation changes (e.g. NADW and Southern Ocean ventilation) in driving climate change across the penultimate and the last deglaciations. Transient simulations performed with Earth-system models that include a dynamic vegetation and a global carbon cycle model, would be particularly useful in assessing the impact of climate change on vegetation cover and on marine ecosystems. The ultimate goal would be to perform transient simulations of the penultimate deglaciation with Earth-system models that include interactive ice sheets and carbon cycle components. But models might not be quite ready for such a task yet.

Transient simulations of the PDG and associated model–paleo-proxy comparisons provide a great opportunity to understand drivers and processes involved in one of the largest natural global warming periods of the Quaternary. In addition, these transient simulations of the PDG will provide a bridge with the proposed PMIP4 transient simulations of the LIG (127 to 121 ka), and the 127 ka time slice experiment (lig127k) (Otto-Bliesner et al.2017). The PDG is a key period to understand as it led to the LIG, an interglacial displaying warmer conditions than the pre-industrial period (e.g. Hoffman et al.2017) as well as a global sea level 6 to 9 m higher than today (e.g. Dutton et al.2015).

Data availability
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Data availability. 

The combined ice-sheet and meltwater scenarios are available in the Supplement. The GHG data can be found at (Köhler et al.2017b). The combined ice sheet as well as the separate Northern Hemispheric and Greenland ice sheets are also publicly available on the Research Data Australia repository at (Menviel et al.2019). In addition, all the forcing files as well as the paleo-data described in the paper are available on the PMIP4 wiki: (last access: May 2019).

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The supplement related to this article is available online at:

Author contributions
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Author contributions. 

LM and EC led and coordinated the study and wrote Sects. 1 and 9. BOB wrote Sect. 2. EC wrote Sect. 3 in collaboration with AL, IO, KK and EW. LT and AAO provided the continental ice-sheet data. LM wrote Sect. 4 in collaboration with LT, AAO, PCT and PLG. AD reassessed the timing of the sea-level changes across the PDG and wrote Sect. 5 in collaboration with EC. LM wrote Sects. 6 and 7 in collaboration with BOB, RFI, LG, FH, MK, LT and XZ. AG and EC in collaboration with RND reassessed all the chronologies of the paleo-records presented in the study. AG, EC, LM and RDN wrote Sect. 8. All the authors designed the goals of the study and contributed to the different parts of the paper.

Competing interests
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Competing interests. 

The authors declare that they have no conflict of interest.

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This study was performed as part of the PAGES-PMIP working group on Quaternary Interglacials (QUIGS) and was initiated during the second QUIGS workshop at the Université du Québec in Montréal, 18–20 October 2016. PMIP is endorsed by the World Climate Research Program (WCRP) and CLIVAR. Andrea Dutton acknowledges the PALSEA working group. We thank J.-Y. Peterschmitt for setting up the PDG page on the PMIP4 wiki. The paper has benefited from helpful comments from two anonymous reviewers as well as from the editor Julia Hargreaves.

Financial support
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Financial support. 

Laurie Menviel acknowledges funding from the Australian Research Council (grant nos. DE150100107 and DP180100048). Emilie Capron received funding from the European Union's Seventh Framework Programme for research and innovation under the Marie Skłodowska-Curie grant agreement no. 600207. Eric Wolff is supported by a Royal Society professorship. His part in this project has received funding from the European Research Council (ERC) under the European Union's Horizon 2020 research and innovation programme (grant agreement no. 742224). Polychronis C. Tzedakis and Ruza F. Ivanovic acknowledge funding from NERC (grant nos. NE/G00756X/1 to Polychronis C. Tzedakis and NE/K008536/1 to Ruza F. Ivanovic). Masa Kageyama is funded by the CNRS. Andrea Dutton acknowledges U.S. National Science Foundation (NSF) (grant nos. 1559040 and 1702740). Bette Otto-Bliesner's contributions are based upon work supported by the National Center for Atmospheric Research, which is a major facility sponsored by the U.S. NSF under Cooperative Agreement no. 1852977, with additional funding provided by an NSF P2C2 grant (grant no. AGS-1401803). Feng He was supported by the NSF (grant no. AGS-1502990) and by the NOAA Climate and Global Change Postdoctoral Fellowship program, administered by the University Corporation for Atmospheric Research. Xu Zhang is supported by Helmholtz Postdoc Program (PD-301) and the Chinese “The Thousand Talents Plan” Program. Ayako Abe-Ouchi, Kenji Kawamura and Ikumi Oyabu acknowledge support by JSPS KAKENHI (grant nos. 17H06104 to Ayako Abe-Ouchi, 15KK0027 and 26241011 to Kenji Kawamura, 17K12816 and 17J00769 to Ikumi Oyabu), and MEXT KAKENHI (grant nos. 17H06323 to Ayako Abe-Ouchi and 17H06320 to Kenji Kawamura).

Review statement
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Review statement. 

This paper was edited by Julia Hargreaves and reviewed by Julia Hargreaves and one anonymous referee.

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Abe-Ouchi, A., Segawa, T., and Saito, F.: Climatic Conditions for modelling the Northern Hemisphere ice sheets throughout the ice age cycle, Clim. Past, 3, 423–438,, 2007. a

Abe-Ouchi, A., Saito, F., Kawamura, K., Raymo, M., Okuno, J., Takahashi, K., and Blatter, H.: Insolation-driven 100,000-year glacial cycles and hysteresis of ice-sheet volume, Nature, 500, 190–193, 2013. a, b, c, d, e, f, g, h, i, j, k, l

Alley, R.: Ice-core evidence of abrupt climate changes, P. Natl. Acad. Sci. USA, 97, 1331–1334, 2000. a

Anderson, R. F., Ali, S., Bradtmiller, L., Nielsen, S., Fleisher, M., Anderson, B., and Burckle, L.: Wind-driven upwelling in the Southern Ocean and the deglacial rise in Atmospheric CO2, Science, 323, 1443–1448, 2009. a

Arz, H., Pätzold, J., and Wefer, G.: Correlated millenial-scale changes in surface hydrography and terrigenous sediment yield inferred from last-glacial marine deposits off Northeastern Brazil, Quaternary Res., 50, 157–166, 1998. a, b

Astakhov, V.: Evidence of Late Pleistocene ice-dammed lakes in West Siberia, Quaternary Sci. Rev., 23, 1285–1311,, 2004. a

Astakhov, V., Shkatova, V., Zastrozhnov, A., and Chuyko, M.: Glaciomorphological Map of the Russian Federation, Quaternary Int., 420, 4–14,, 2016. a

Austermann, J., Mitrovica, J., Latychev, K., and Milne, G.: Barbados-based estimate of ice volume at Last Glacial Maximum affected by subducted plate, Nat. Geosci., 6, 553–557,, 2013. a

Bakker, P., Stone, E. J., Charbit, S., Gröger, M., Krebs-Kanzow, U., Ritz, S. P., Varma, V., Khon, V., Lunt, D. J., Mikolajewicz, U., Prange, M., Renssen, H., Schneider, B., and Schulz, M.: Last interglacial temperature evolution – a model inter-comparison, Clim. Past, 9, 605–619,, 2013. a

Balaji, V., Taylor, K. E., Juckes, M., Lawrence, B. N., Durack, P. J., Lautenschlager, M., Blanton, C., Cinquini, L., Denvil, S., Elkington, M., Guglielmo, F., Guilyardi, E., Hassell, D., Kharin, S., Kindermann, S., Nikonov, S., Radhakrishnan, A., Stockhause, M., Weigel, T., and Williams, D.: Requirements for a global data infrastructure in support of CMIP6, Geosci. Model Dev., 11, 3659–3680,, 2018. a

Bard, E., Hamelin, B., and Fairbanks, R.: U-Th ages obtained by mass spectrometry in corals from Barbados: sea level during the past 130,000 years, Nature, 346, 456–458,, 1990. a, b

Bard, E., Hamelin, B., and Delanghe-Sabatier, D.: Deglacial Meltwater Pulse 1B and Younger Dryas Sea Levels Revisited with Boreholes at Tahiti, Science, 327, 1235–1237, 2010. a

Barker, S., Diz, P., Vautravers, M., Pike, J., Knorr, G., Hall, I., and Broecker, W.: Interhemispheric Atlantic seesaw response during the last deglaciation, Nature, 457, 1097–1102, 2009. a

Barker, S., Knorr, G., Edwards, R. L., Parrenin, F., Putnam, A. E., Skinner, L. C., Wolff, E., and Ziegler, M.: 800,000 Years of Abrupt Climate Variability, Science, 334, 347–351,, 2011. a, b, c

Barker, S., Chen, J., Gong, X., Jonkers, L., Knorr, G., and Thornalley, D.: Icebergs not the trigger for North Atlantic cold events, Nature, 520, 333–336, 2015. a, b, c

Baumgartner, M., Schilt, A., Eicher, O., Schmitt, J., Schwander, J., Spahni, R., Fischer, H., and Stocker, T. F.: High-resolution interpolar difference of atmospheric methane around the Last Glacial Maximum, Biogeosciences, 9, 3961–3977,, 2012. a

Bazin, L., Landais, A., Lemieux-Dudon, B., Toyé Mahamadou Kele, H., Veres, D., Parrenin, F., Martinerie, P., Ritz, C., Capron, E., Lipenkov, V., Loutre, M.-F., Raynaud, D., Vinther, B., Svensson, A., Rasmussen, S. O., Severi, M., Blunier, T., Leuenberger, M., Fischer, H., Masson-Delmotte, V., Chappellaz, J., and Wolff, E.: An optimized multi-proxy, multi-site Antarctic ice and gas orbital chronology (AICC2012): 120–800 ka, Clim. Past, 9, 1715–1731,, 2013. a, b, c, d, e, f

Bereiter, B., Eggleston, S., Schmitt, J., Nehrbass-Ahles, C., Stocker, T., Fischer, H., Kipfstuhl, S., and Chappellaz, J.: Revision of the EPICA Dome C CO2 record from 800 to 600 kyr before present, Geophys. Res. Lett., 42, 542–549,, 2015. a, b

Berger, A.: Long term variations of daily insolation and Quaternary climate change, J. Atmos. Sci., 35, 2362–2367, 1978. a, b, c, d, e, f

Blunier, T. and Brook, E.: Timing of millennial-scale climate change in Antarctica and Greenland during the last glacial period, Science, 291, 109–112, 2001. a

Böhm, E., Lippold, J., Gutjahr, M., Frank, M., Blaser, P., Antz, B., Fohlmeister, J., Frank, N., Andersen, M., and Deininger, M.: Strong and deep Atlantic meridional overturning circulation during the last glacial cycle, Nature, 517, 73–76, 2015. a, b, c, d, e, f

Bond, G., Heinrich, H., Broecker, W., and Labeyrie, L.: Evidence of massive discharges of icebergs into the North Atlantic during the last glacial period, Nature, 360, 245–249, 1992. a

Bopp, L., Kohfeld, K., Quéré, C. L., and Aumont, O.: Dust impact on marine biota and atmospheric CO2 during glacial periods, Paleoceanography, 18, 1046,, 2003. a

Bradley, S. L., Reerink, T. J., van de Wal, R. S. W., and Helsen, M. M.: Simulation of the Greenland Ice Sheet over two glacial-interglacial cycles: investigating a sub-ice- shelf melt parameterization and relative sea level forcing in an ice-sheet-ice-shelf model, Clim. Past, 14, 619–635,, 2018. a

Brauer, A., Allen, J., Mingram, J. J., Dulski, P., Wulf, S., and Huntley, B.: Evidence for last interglacial chronology and environmental change from Southern Europe, P. Natl. Acad. Sci. USA, 104, 450–455, 2007. a, b

Briggs, R., Pollard, D., and Tarasov, L.: A glacial systems model configured for large ensemble analysis of Antarctic deglaciation, The Cryosphere, 7, 1949–1970,, 2013. a, b

Briggs, R., Pollard, D., and Tarasov, L.: A data-constrained large ensemble analysis of Antarctic evolution since the Eemian, Quaternary Sci. Rev., 103, 91–115,, 2014. a, b, c, d, e, f, g, h, i, j

Buiron, D., Chappellaz, J., Stenni, B., Frezzotti, M., Baumgartner, M., Capron, E., Landais, A., Lemieux-Dudon, B., Masson-Delmotte, V., Montagnat, M., Parrenin, F., and Schilt, A.: TALDICE-1 age scale of the Talos Dome deep ice core, East Antarctica, Clim. Past, 7, 1–16,, 2011. a

Buizert, C., Gkinis, V., Severinghaus, J., He, F., Lecavalier, B., Kindler, P., Leuenberger, M., Carlson, A., Vinther, B., Masson-Delmotte, V., White, J., Liu, Z., Otto-Bliesner, B., and Brook, E.: Greenland temperature response to climate forcing during the last deglaciation, Science, 345, 1177–1180, 2014. a, b, c, d

Buizert, C., Cuffey, K. M., Severinghaus, J. P., Baggenstos, D., Fudge, T. J., Steig, E. J., Markle, B. R., Winstrup, M., Rhodes, R. H., Brook, E. J., Sowers, T. A., Clow, G. D., Cheng, H., Edwards, R. L., Sigl, M., McConnell, J. R., and Taylor, K. C.: The WAIS Divide deep ice core WD2014 chronology – Part 1: Methane synchronization (68–31 ka BP) and the gas age–ice age difference, Clim. Past, 11, 153–173,, 2015. a

Caley, T., Roche, D., and Renssen, H.: Orbital Asian summer monsoon dynamics revealed using an isotope-enabled global climate model, Nat. Commun., 5, 5371,, 2014. a

Capron, E., Landais, A., Chappellaz, J., Schilt, A., Buiron, D., Dahl-Jensen, D., Johnsen, S. J., Jouzel, J., Lemieux-Dudon, B., Loulergue, L., Leuenberger, M., Masson-Delmotte, V., Meyer, H., Oerter, H., and Stenni, B.: Millennial and sub-millennial scale climatic variations recorded in polar ice cores over the last glacial period, Clim. Past, 6, 345–365,, 2010. a

Capron, E., Govin, A., Stone, E., Masson-Delmotte, V., Mulitza, S., Otto-Bliesner, B., Rasmussen, T., Sime, L., Waelbroeck, C., and Wolff, E.: Temporal and spatial structure of multi-millennial temperature changes at high latitudes during the Last Interglacial, Quaternary Sci. Rev., 103, 116–133, 2014. a

Capron, E., Govin, A., Feng, R., Otto-Bliesner, B., and Wolff, E.: Critical evaluation of climate syntheses to benchmark CMIP6/PMIP4 127 ka Last Interglacial simulations in the high-latitude regions, Quaternary Sci. Revi., 168, 137–150,, 2017. a

Carlson, A. and Winsor, K.: Northern Hemisphere ice-sheet responses to past climate warming, Nat. Geosci., 5, 507–613,, 2012. a, b

Chapman, M. and Shackleton, N.: Millennial-scale fluctuations in North Atlantic heat flux during the last 150,000 years, Earth Planet. Sci. Lett., 159, 57–70, 1998. a, b, c, d, e, f

Chapman, M., Shackleton, N., and Duplessy, J.-C.: Sea surface temperature variability during the last glacial-interglacial cycle: assessing the magnitude and pattern of climate change in the North Atlantic, Palaeogeogr. Palaeoclimatol. Palaeoecol., 157, 1–25, 2000. a

Cheng, H., Edwards, R. L., Broecker, W. S., Denton, G. H., Kong, X., Wang, Y., Zhang, R., and Wang, X.: Ice Age Terminations, Science, 326, 248–252,, 2009. a, b, c, d, e, f, g, h, i

Cheng, H., Edwards, R. L., Shen, C.-C., Polyak, V. J., Asmerom, Y., Woodhead, J., Hellstrom, J., Wang, Y., Kong, X., Spötl, C., Wang, X., and Alexander, E. C.: Improvements in 230Th dating, 230Th and 234U half-life values, and U-Th isotopic measurements by multi-collector inductively coupled plasma mass spectrometry, Earth Planet. Sci. Lett., 371–372, 82–91,, 2013. a, b

Cheng, H., Edwards, R., Sinha, A., Spötl, C., Yi, L., Chen, S., Kelly, M., Kathayat, G., Wang, X., Li, X., Kong, X., Wang, Y., Ning, Y., and Zhang, H.: The Asian monsoon over the past 640,000 years and ice age terminations, Nature, 534, 640–646,, 2016. a, b, c, d

Chiang, J. and Bitz, C.: Influence of high latitude ice cover on the marine Intertropical Convergence Zone, Clim. Dynam., 25, 477–496, 2005. a, b

Colleoni, F., Wekerle, C., Näslund, J.-O., Brandefelt, J., and Masina, S.: Constraint on the penultimate glacial maximum Northern Hemisphere ice topography (∼140 kyrs BP), Quaternary Sci. Rev., 137, 97–112, 2016. a

Cortijo, E., Lehman, S., Keigwin, L., Chapman, M., Paillard, D., and Labeyrie, L.: Changes in Meridional Temperature and Salinity Gradients in the North Atlantic Ocean (30–72 N) during the Last Interglacial Period, Paleoceanography, 14, 23–33,, 1999. a

Curry, B., Grimley, D., and McKay, E.: Quaternary Glaciations: Extent and Chronology – A Closer Look, vol. 15 of Developments in Quaternary Science, chap. Quaternary glaciations in Illinois, 467–487, Elsevier, 2011. a, b

Curry, W., Duplessy, J., Labeyrie, L., and Shackleton, N.: Changes in the distribution of δ13C of deep water CO2 between the last glaciation and the Holocene, Paleoceanography, 3, 317–341,, 1988. a

Deaney, E., Barker, S., and van de Flierdt, T.: Timing and nature of AMOC recovery across Termination 2 and magnitude of deglacial CO2 change, Nat. Commun., 8, 14595,, 2017. a, b, c, d, e, f, g, h, i, j

Denton, G., Anderson, R., Toggweiler, J., Edwards, R., Schaefer, J., and Putnam, A.: The last glacial termination, Science, 328, 1652–1656, 2010. a

Deplazes, G., Lückge, A., Peterson, L., Timmermann, A., Hamann, Y., Hughen, K., Röh, U., Laj, C., Cane, M., Sigman, D., and Haug, G.: Links between tropical rainfall and North Atlantic climate during the last glacial period, Nat. Geosci., 6, 213–217,, 2013. a, b

Deschamps, P., Durand, N., Bard, E., Hamelin, B., Camoin, G., Thomas, A., Henderson, G., Okuno, J., and Yokohama, Y.: Ice-sheet collapse and sea-level rise at the Bølling warming 14,600 years ago, Nature, 483, 559–564,, 2012. a

Drysdale, R., Zanchetta, G., Hellstrom, J., Fallick, A., McDonald, J., and Cartwright, I.: Stalagmite evidence for the precise timing of North Atlantic cold events during the early last glacial, Geology, 35, 77–80, 2007. a

Drysdale, R., Hellstrom, J., Zanchetta, G., Fallick, A., Sanchez-Goni, M., Couchoud, I., McDonald, J., Maas, R., Lohmann, G., and I.Isola: Evidence for Obliquity Forcing of Glacial Termination II, Science, 325, 1527–1531, 2009. a, b, c

Drysdale, R., Couchoud, I., Zanchetta, G., Isola, I., Regattieri, E., Hellstrom, J., Tzedakis, P., Ireland, T., Greig, G., Wong, H., Govin, A., Corrick, E., Holden, P., and Woodhead, J.: Magnesium as a paleotemperature proxy in a slow-growing subaqueous speleothem, Geology, in preparation, 2019. a, b

Duplessy, J., Shackleton, N., Fairbanks, R., Labeyrie, L., Oppo, D., and Kallel, N.: Deepwater source variations during the last climate cycle and their impact on the global deepwater circulation, Paleoceanography, 3, 343–360,, 1988. a

Dutton, A. and Lambeck, K.: Ice Volume and Sea Level During the Last Interglacial, Science, 337, 216–219,, 2012. a

Dutton, A., Carlson, A., Long, A., Milne, G., Clark, P., DeConto, R., Horton, B., Rahmstorf, S., and Raymo, M.: Sea-level rise due to polar ice-sheet mass loss during past warm periods, Science, 349, aaa4019-1,, 2015. a, b, c, d, e

Dyke, A.: Quaternary Glaciations – Extent and Chronology – Part II: North America, Developments in Quaternary Sciences, chap. An outline of North American deglaciation with emphasis on central and northern Canada, 373–424, Elsevier, Amsterdam, the Netherlands, 2004. a

Dyke, A., Andrews, J., Clark, P., England, J., Miller, G., Shaw, J., and Veillette, J.: The Laurentide and Innuitian ice sheets during the Last Glacial Maximum, Quaternary Sci. Rev., 21, 9–31, 2002. a, b

Ehlers, J., Grube, A., Stephan, H.-J., and Wansa, S.: Quaternary Glaciations: Extent and Chronology – A Closer Look, vol. 15 of Developments in Quaternary Science, chap. Pleistocene glaciations of North Germany – New results, 149–162, elsevier edn., 2011. a, b

EPICA community members: One-to-one coupling of glacial climate variability in Greenland and Antarctica, Nature, 444, 195–198, 2006. a

Esat, T. M., McCulloch, M. T., Chappell, J., Pillans, B., and Omura, A.: Rapid Fluctuations in Sea Level Recorded at Huon Peninsula During the Penultimate Deglaciation, Science, 283, 197–201,, 1999. a, b, c, d, e

Evan, A. T., Vimont, D. J., Heidinger, A. K., Kossin, J. P., and Bennartz, R.: The Role of Aerosols in the Evolution of Tropical North Atlantic Ocean Temperature Anomalies, Science, 324, 778–781,, 2009. a

Flückiger, J., Monnin, E., Stauffer, B., Schwander, J., Stocker, T., Chappellaz, J., Raynaud, D., and Barnola, J.-M.: High-resolution Holocene N2O ice core record and its relationship with CH4 and CO2, Global Biogeochem. Cy., 16, 10-1–10-8,, 2002. a, b

Fujita, K., Omori, A., Yokoyama, Y., Sakai, S., and Iryu, Y.: Sea-level rise during Termination II inferred from large benthic foraminifers: IODP Expedition 310, Tahiti Sea Level, Mar. Geol., 271, 149–155,, 2010. a

Gherardi, J., Labeyrie, L., Nave, S., Francois, R., McManus, J., and Cortijo, E.: Glacial-interglacial circulation changes inferred from 231Pa/230Th sedimentary record in the North Atlantic region, Paleoceanography, 24, PA2204,, 2009. a

Goelzer, H., Huybrechts, P., Loutre, M.-F., and Fichefet, T.: Last Interglacial climate and sea-level evolution from a coupled ice sheet–climate model, Clim. Past, 12, 2195–2213,, 2016a. a, b, c, d, e, f, g

Goelzer, H., Huybrechts, P., Loutre, M.-F., and Fichefet, T.: Impact of ice sheet meltwater fluxes on the climate evolution at the onset of the Last Interglacial, Clim. Past, 12, 1721–1737,, 2016b. a

Golledge, N., L. Menviel and, L. C., Fogwill, C., England, M., Cortese, G., and Levy, R.: Antarctic contribution to meltwater pulse 1A from reduced Southern Ocean overturning, Nat. Commun., 5, 5107,, 2014. a

Gong, X., Zhang, X., Lohmann, G., Wei, W., Zhang, X., and Pfeiffer, M.: Higher Laurentide and Greenland ice sheets strengthen the North Atlantic ocean circulation, Clim. Dynam., 45, 139–150,, 2015. a, b

Govin, A., Michel, E., Labeyrie, L., Waelbroeck, C., Dewilde, F., and Jansen, E.: Evidence for northward expansion of Antarctic Bottom Water mass in the Southern Ocean during the last glacial inception, Paleoceanography, 24, PA1202,, 2009. a, b

Govin, A., Braconnot, P., Capron, E., Cortijo, E., Duplessy, J.-C., Jansen, E., Labeyrie, L., Landais, A., Marti, O., Michel, E., Mosquet, E., Risebrobakken, B., Swingedouw, D., and Waelbroeck, C.: Persistent influence of ice sheet melting on high northern latitude climate during the early Last Interglacial, Clim. Past, 8, 483–507,, 2012. a, b, c, d, e

Govin, A., Capron, E., Tzedakis, P., Verheyden, S., Ghaleb, B., Hillaire-Marcel, C., St-Onge, G., Stoner, J., Bassinot, F., Bazin, L., Blunier, T., Combourieu-Nebout, N., Ouahabi, A. E., Genty, D., Gersonde, R., Jimenez-Amat, P., Landais, A., Martrat, B., Masson-Delmotte, V., Parrenin, F., Seidenkrantz, M.-S., Veres, D., Waelbroeck, C., and Zahn, R.: Sequence of events from the onset to the demise of the Last Interglacial: Evaluating strengths and limitations of chronologies used in climatic archives, Quaternary Sci. Rev., 129, 1–36, 2015. a, b, c, d, e, f, g, h, i

Grant, K., Rohling, E., Bar-Matthews, M., Ayalon, A., Medina-Elizalde, M., Ramsey, C. B., Satow, C., and Roberts, A.: Rapid coupling between ice volume and polar temperature over the past 150,000 years, Nature, 491, 744–747,, 2012. a, b, c, d, e, f, g, h, i, j, k, l, m, n, o

Grant, K., Rohling, E., Ramsey, C. B., Cheng, H., Edwards, R., Florindo, F., Heslop, D., Marra, F., Roberts, A., Tamisiea, M., and Williams, F.: Sea-level variability over five glacial cycles, Nat. Commun., 5, 5076,, 2014. a, b, c, d, e, f

Gregoire, L., Payne, A., and Valdes, P.: Deglacial rapid sea level rises caused by ice-sheet saddle collapses, Nature, 487, 219–222,, 2012. a, b, c

Gregoire, L., Valdes, P., and Payne, A.: The relative contribution of orbital forcing and greenhouse gases to the North American deglaciation, Geophys. Res. Lett., 42, 9970–9979,, 2015. a

Gregoire, L., Otto-Bliesner, B., Valdes, P., and Ivanovic, R.: Abrupt Bølling warming and ice saddle collapse contributions to the Meltwater Pulse 1a rapid sea level rise, Geophys. Res. Lett., 43, 9130–9137,, 2016. a

Gregoire, L., Ivanovic, R., Maycock, A., Valdes, P., and Stevenson, S.: Holocene lowering of the Laurentide ice sheet affects North Atlantic gyre circulation and climate, Clim. Dynam., 51, 3797–3813,, 2018. a

He, F., Shakun, J., Clark, P., Carlson, A., Liu, Z., Otto-Bliesner, B., and Kutzbach, J.: Northern Hemisphere forcing of Southern Hemisphere climate during the last deglaciation, Nature, 494, 81–85, 2013. a, b, c, d, e

Head, M. J. and Gibbard, P. L.: Formal subdivision of the Quaternary System/Period: Past, present, and future, Quaternary Int., 383, 4–35,, 2015. a

Heinrich, H.: Origin and consequences of cyclic ice rafting in the northeast Atlantic Ocean during the past 130,000 years, Quaternary Res., 29, 142–152, 1988. a, b

Hibbert, F. D., Rohling, E. J., Dutton, A., Williams, F. H., Chutcharavan, P. M., Zhao, C., and Tamisiea, M. E.: Coral indicators of past sea-level change: A global repository of U-series dated benchmarks, Quaternary Sci. Rev., 145, 1–56,, 2016. a

Hodell, D., Anselmetti, F., Ariztegui, D., Brenner, M., Curtis, J., Gilli, A., Grzesik, D., Guilderson, T., Müller, A., Bush, M., Correa-Metrio, A., Escobar, J., and Kutterolf, S.: An 85-ka record of climate change in lowland Central America, Quaternary Sci. Rev., 27, 1152–1165, 2008. a, b, c

Hodell, D., Nicholl, J., Bontognali, T., Danino, S., Dorador, J., Dowdeswell, J., Einsle, J., Kuhlmann, H., Martrat, B., Mleneck-Vautravers, M., Rodriguez-Tovar, F., and Röhl, U.: Anatomy of Heinrich Layer 1 and its role in the last deglaciation, Paleoceanography, 32, 284–303,, 2017. a

Hodell, D. A., Minth, E. K., Curtis, J. H., McCave, I. N., Hall, I. R., Channell, J. E., and Xuan, C.: Surface and deep-water hydrography on Gardar Drift (Iceland Basin) during the last interglacial period, Earth Planet. Sci. Lett., 288, 10–19,, 2009. a

Hoffman, J., Clark, P., Parnell, A., and He, F.: Regional and global sea-surface temperatures during the last interglaciation, Science, 355, 276–279, 2017. a, b, c, d, e

Hu, A., Meehl, G., Han, W., Timmermann, A., Otto-Bliesner, B., Liu, Z., Washington, W., Large, W., Abe-Ouchi, A., Kimoto, M., Lambeck, K., and Wu, B.: Role of the Bering Strait on the hysteresis of the ocean conveyor belt circulation and glacial climate stability, P. Natl. Acad. Sci. USA, 107, 6417–6422,, 2012. a

Hughes, A., Gyllencreutz, R., Lohne, O., Mangerud, J., and Svendsen, J.: The last Eurasian ice sheets – a chronological database and time-slice reconstruction, DATED-1, Boreas, 45, 1–45,, 2016. a

Hughes, P. and Gibbard, P.: Global glacier dynamics during 100 ka Pleistocene glacial cycles, Quaternary Res., 90, 222–243,, 2019. a, b, c

Hut, G.: Stable isotope reference samples for geochemical and hydrological investigations, Tech. rep., International Atomic Energy Agency, Vienna 42, 1987. a

Ivanovic, R. F., Gregoire, L. J., Kageyama, M., Roche, D. M., Valdes, P. J., Burke, A., Drummond, R., Peltier, W. R., and Tarasov, L.: Transient climate simulations of the deglaciation 21-9 thousand years before present (version 1) – PMIP4 Core experiment design and boundary conditions, Geosci. Model Dev., 9, 2563–2587,, 2016. a, b, c, d, e, f, g, h

Ivanovic, R., Gregoire, L., Wickert, A., Valdes, P., and Burke, A.: Collapse of the North American ice saddle 14,500 years ago caused widespread cooling and reduced ocean overturning circulation, Geophys. Res. Lett., 44, 383–392,, 2017. a

Ivanovic, R., Gregoire, L., Burke, A., Wickert, A., Valdes, P., Ng, H., Robinson, L., McManus, J., Mitrovica, J., Lee, L., and Dentith, J.: Acceleration of northern ice sheet melt induces AMOC slowdown and northern cooling in simulations of the early last deglaciation, Paleoceanogr. Paleoclimatol., 33, 807–824,, 2018. a, b

Jacobel, A. W., McManus, J. F., Anderson, R. F., and Winckler, G.: Large deglacial shifts of the Pacific Intertropical Convergence Zone, Nat. Commun., 7, 10449,, 2016. a

Jonkers, L. and Kučera, M.: Global analysis of seasonality in the shell flux of extant planktonic Foraminifera, Biogeosciences, 12, 2207–2226,, 2015. a

Jouzel, J., Vaikmae, R., Petit, J., Martin, M., Duclos, Y., Stievenard, M., Lorius, C., Toots, M., Melieres, M., Burckle, L., Barkov, N., and Kotlyakov, V.: The two-steps shape and timing of the last deglaciation in Antarctica, Clim. Dynam., 11, 151–161, 1995. a

Jouzel, J., Masson, V., Cattani, O., Dreyfus, G., Falourd, S., Hoffmann, G., Minster, B., Nouet, J., Barnola, J., Chappellaz, J., Fischer, H., Gallet, J., Leuenberger, S. J. M., , Loulergue, L., Luethi, D., Oerter, H., Parrenin, F., Raisbeck, G., Raynaud, D., Schilt, A., Schwander, J., Selmo, E., Souchez, R., Spahni, R., Stauffer, B., Steffenssen, J. P., Stenni, B., Stocker, T., Tison, J., Werner, M., and Wolff, E.: Orbital and millennial Antarctic climate variability over the past 800,000 years, Science, 317, 793–796, 2007. a, b, c, d, e, f

Kageyama, M., Mignot, J., Swingedouw, D., Marzin, C., Alkama, R., and Marti, O.: Glacial climate sensitivity to different states of the Atlantic Meridional Overturning Circulation: results from the IPSL model, Clim. Past, 5, 551–570,, 2009. a

Kageyama, M., Paul, A., Roche, D., and Meerbeeck, C. V.: Modelling glacial climatic millennial-scale variability related to changes in the Atlantic meridional overturning circulation: a review, Quaternary Sci. Rev., 29, 2931–2956,, 2010. a

Kageyama, M., Merkel, U., Otto-Bliesner, B., Prange, M., Abe-Ouchi, A., Lohmann, G., Ohgaito, R., Roche, D. M., Singarayer, J., Swingedouw, D., and Zhang, X.: Climatic impacts of fresh water hosing under Last Glacial Maximum conditions: a multi-model study, Clim. Past, 9, 935–953,, 2013. a, b, c

Kageyama, M., Albani, S., Braconnot, P., Harrison, S. P., Hopcroft, P. O., Ivanovic, R. F., Lambert, F., Marti, O., Peltier, W. R., Peterschmitt, J.-Y., Roche, D. M., Tarasov, L., Zhang, X., Brady, E. C., Haywood, A. M., LeGrande, A. N., Lunt, D. J., Mahowald, N. M., Mikolajewicz, U., Nisancioglu, K. H., Otto-Bliesner, B. L., Renssen, H., Tomas, R. A., Zhang, Q., Abe-Ouchi, A., Bartlein, P. J., Cao, J., Li, Q., Lohmann, G., Ohgaito, R., Shi, X., Volodin, E., Yoshida, K., Zhang, X., and Zheng, W.: The PMIP4 contribution to CMIP6 – Part 4: Scientific objectives and experimental design of the PMIP4-CMIP6 Last Glacial Maximum experiments and PMIP4 sensitivity experiments, Geosci. Model Dev., 10, 4035–4055,, 2017. a, b, c, d, e

Keigwin, L., Klotsko, S., Zhao, N., Reilly, B., Giosan, L., and Driscoll, N.: Deglacial floods in the Beaufort Sea preceded Younger Dryas cooling, Nat. Geosci., 11, 599–604,, 2018. a

Kohfeld, K. and Ridgwell, A.: Surface Ocean-Lower Atmosphere Processes, chap. Glacial-interglacial variability in atmospheric CO2, Geophysical Research Series 187, American Geophysical Union, 2009. a

Köhler, P., Nehrbass-Ahles, C., Schmitt, J., Stocker, T. F., and Fischer, H.: A 156 kyr smoothed history of the atmospheric greenhouse gases CO2, CH4, and N2O and their radiative forcing, Earth Syst. Sci. Data, 9, 363–387,, 2017a. a, b, c, d, e, f, g, h, i, j, k, l

Köhler, P., Nehrbass-Ahles, C., Schmitt, J., Stocker, T. F., and Fischer, H.: Compilations and splined-smoothed calculations of continuous records of the atmospheric greenhouse gases CO2, CH4, and N2O and their radiative forcing since the penultimate glacial maximum,, 2017b. a

Kopp, R., Simons, F., Mitrovica, J., Maloof, A., and Oppenheimer, M.: Probabilistic assessment of sea level during the last interglacial stage, Nature, 462, 863–867,, 2009. a, b

Labeyrie, L., Leclaire, H., Waelbroeck, C., Cortijo, E., Duplessy, J., Vidal, L., Elliot, M., Lecoat, B., and Auffret, G.: Mechanisms of Global Climate Change at Millennial Time Scales, vol. 112, chap. Insolation forcing and millennial scale variability of the North WestAtlantic Ocean: Surface versus deep water changes, 77–98, Geophy. Monogr. Series,AGU, Washington, D. C., 1999. a, b, c, d, e, f, g

Lambeck, K., Purcell, A., Funder, S., Kjaer, K., Larsen, E., and Möller, P.: Constraints on the Late Saalian to early Middle Weichselian ice sheet of Eurasia from field data and rebound modelling, Boreas, 35, 539–575, 2006. a, b, c

Lambeck, K., Rouby, H., Purcell, A., Sun, Y., and Sambridge, M.: Sea level and global ice volumes from the Last Glacial Maximum to the Holocene, P. Natl. Acad. Sci. USA, 111, 15296–15303,, 2014. a, b, c, d

Landais, A., Dreyfus, G., Capron, E., Jouzel, J., Masson-Delmotte, V., Roche, D. M., Prié, F., Caillon, N., Chappellaz, J., Leuenberger, M., Lourantou, A., Parrenin, F., Raynaud, D., and Teste, G.: Two-phase change in CO2, Antarctic temperature and global climate during Termination II, Nat. Geosci., 6, 1062–1065, 2013. a, b, c, d, e, f, g

Landais, A., Masson-Delmotte, V., Capron, E., Langebroek, P. M., Bakker, P., Stone, E. J., Merz, N., Raible, C. C., Fischer, H., Orsi, A., Prié, F., Vinther, B., and Dahl-Jensen, D.: How warm was Greenland during the last interglacial period?, Clim. Past, 12, 1933–1948,, 2016. a

Le Morzadec, K., Tarasov, L., Morlighem, M., and Seroussi, H.: A new sub-grid surface mass balance and flux model for continental-scale ice sheet modelling: testing and last glacial cycle, Geosci. Model Dev., 8, 3199–3213,, 2015. a

Lisiecki, L. E. and Raymo, M. E.: A Pliocene-Pleistocene stack of 57 globally distributed benthic δ18O records, Paleoceanography, 20, PA1003,, 2005. a, b, c, d, e

Liu, Z., Otto-Bliesner, B., He, F., Brady, E., Tomas, R., Clark, P., Carlson, A., Lynch-Stieglitz, J., Curry, W., Brook, E., Erickson, D., Jacob, R., Kutzbach, J., and Cheng, J.: Transient simulation of last deglaciation with a new mechanism for Bølling-Allerød warming, Science, 325, 310–314, 2009. a, b, c, d, e

Löfverström, M., Caballero, R., Nilsson, J., and Kleman, J.: Evolution of the large-scale atmospheric circulation in response to changing ice sheets over the last glacial cycle, Clim. Past, 10, 1453–1471,, 2014. a

Loulergue, L., Schilt, A., Spahni, R., Masson-Delmotte, V., Blunier, T., Lemieux, B., Barnola, J.-M., Raynaud, D., Stocker, T., and Chappellaz, J.: Orbital and millennial-scale features of atmospheric CH4 over the past 800,000 years, Nature, 453, 383–386, 2008. a, b, c, d, e

Lourantou, A., Lavric, J., Kohler, P., Barnola, J.-M., Paillard, D., Michel, E., Raynaud, D., and Chappellaz, J.: Constraint of the CO2 rise by new atmospheric carbon isotopic measurements during the last deglaciation, Global Biogeochem. Cy., 24, GB2015,, 2010. a, b, c, d

Loutre, M. F., Fichefet, T., Goosse, H., Huybrechts, P., Goelzer, H., and Capron, E.: Factors controlling the last interglacial climate as simulated by LOVECLIM1.3, Clim. Past, 10, 1541–1565,, 2014. a, b, c, d

Lüthi, D., Floch, M. L., Bereiter, B., Blunier, T., Barnola, J.-M., Siegenthaler, U., Raynaud, D., Jouzel, J., Fischer, H., Kawamura, K., and Stocker, T.: High-resolution carbon dioxide concentration record 650,000–800,000 years before present, Nature, 453, 379–382,, 2008. a, b

Marchal, O., T.F.Stocker, and F.Joos: A latitude-depth, circulation-biogeochemical ocean model for paleoclimate studies. Development and sensitivities, Tellus B, 50, 290–316,, 1998. a

Marcott, S., Bauska, T., Buizert, C., Steig, E., Rosen, J., Cuffey, K., Fudge, T., Severinghaus, J., Ahn, J., Kalk, M., McConnell, J., Sowers, T., Taylor, K., White, J., and Brook, E.: Centennial-scale changes in the global carbon cycle during the last deglaciation, Nature, 514, 616–619,, 2014. a, b, c, d

Margari, V., Skinner, L. C., Tzedakis, P. C., Ganopolski, A., Vautravers, M., and Shackleton, N. J.: The nature of millennial-scale climate variability during the past two glacial periods, Nat. Geosci., 3, 127–131, 2010. a, b

Margari, V., Skinner, L. C., Hodell, D. A., Martrat, B., Toucanne, S., Grimalt, J. O., Gibbard, P. L., Lunkka, J., and Tzedakis, P.: Land-ocean changes on orbital and millennial time scales and the penultimate glaciation, Geology, 42, 183,, 2014. a, b

MARGO project members: Constraints on the magnitude and patterns of ocean cooling at the Last Glacial Maximum, Nat. Geosci., 2, 127–132, 2009. a

Marino, G., Rohling, E., Rodriguez-Sanz, L., Grant, K., Heslop, D., Roberts, A., Stanford, J., and Yu, J.: Bipolar seesaw control on last interglacial sea level, Nature, 522, 197–204, 2015. a, b, c, d

Martin, J.: Glacial-interglacial CO2 change: The iron hypothesis, Paleoceanography, 5, 1–13, 1990. a

Martínez-García, A., Sigman, D., Ren, H., Anderson, R., Straub, M., Hodell, D., Jaccard, S., Eglinton, T., and Haug, G.: Iron Fertilization of the Subantarctic Ocean During the Last Ice Age, Science, 343, 1347–1350, 2014. a, b

Martrat, B., Grimalt, J., Shackleton, N., de Abreu, L., Hutterli, M., and Stocker, T.: Four climate cycles of recurring deep and surface water destabilizations on the Iberian margin, Science, 317, 502–507, 2007. a, b, c

Martrat, B., Jimenez-Amat, P., Zahn, R., and Grimalt, J.: Similarities and dissimilarities between the last two deglaciations and interglaciations in the North Atlantic region, Quaternary Sci. Rev., 99, 122–134, 2014. a, b, c, d, e, f, g, h, i, j, k, l, m, n

Marzin, C., Kallel, N., Kageyama, M., Duplessy, J.-C., and Braconnot, P.: Glacial fluctuations of the Indian monsoon and their relationship with North Atlantic climate: new data and modelling experiments, Clim. Past, 9, 2135–2151,, 2013. a

Marzocchi, A. and Jansen, M. F.: Connecting Antarctic sea ice to deep-ocean circulation in modern and glacial climate simulations, Geophys. Res. Lett., 44, 6286–6295,, 2017. a

Masson-Delmotte, V., Buiron, D., Ekaykin, A., Frezzotti, M., Gallée, H., Jouzel, J., Krinner, G., Landais, A., Motoyama, H., Oerter, H., Pol, K., Pollard, D., Ritz, C., Schlosser, E., Sime, L. C., Sodemann, H., Stenni, B., Uemura, R., and Vimeux, F.: A comparison of the present and last interglacial periods in six Antarctic ice cores, Clim. Past, 7, 397–423,, 2011. a, b, c

McGee, D., Donohoe, A., Marshall, J., and Ferreira, D.: Changes in ITCZ location and cross-equatorial heat transport at the Last Glacial Maximum, Heinrich Stadial 1, and the mid-Holocene, Earth Planet. Sci. Lett., 390, 69–79,, 2014. a

McManus, J. F., Oppo, D. W., and Cullen, J. L.: A 0.5-Million-Year Record of Millennial-Scale Climate Variability in the North Atlantic, Science, 283, 971–975,, 1999. a

McManus, J. F., Francois, R., Gherardi, J. M., Keigwin, L. D., and Brown-Leger, S.: Collapse and rapid resumption of Atlantic meridional circulation linked to deglacial climate changes, Nature, 428, 834–837, 2004. a, b, c

Menviel, L., Timmermann, A., Mouchet, A., and Timm, O.: Climate and biogeochemical response to a rapid melting of the West-Antarctic Ice Sheet during interglacials and implications for future climate, Paleoceanography, 25, PA4231,, 2010. a

Menviel, L., Timmermann, A., Timm, O., and Mouchet, A.: Deconstructing the last Glacial Termination: the role of millennial and orbital-scale forcings, Quaternary Sci. Rev., 30, 1155–1172, 2011. a, b, c, d, e

Menviel, L., England, M., Meissner, K., Mouchet, A., and Yu, J.: Atlantic-Pacific seesaw and its role in outgassing CO2 during Heinrich events, Paleoceanography, 29, 58–70,, 2014. a, b

Menviel, L., Mouchet, A., Meissner, K., Joos, F., and England, M.: Impact of oceanic circulation changes on atmospheric δ13CO2, Global Biogeochem. Cy., 29, 1944–1961,, 2015. a

Menviel, L., Yu, J., Joos, F., Mouchet, A., Meissner, K., and England, M.: Poorly ventilated deep ocean at the Last Glacial Maximum inferred from carbon isotopes: a data-model comparison study, Paleoceanography, 32, 2–17,, 2017. a

Menviel, L., Spence, P., Yu, J., Chamberlain, M., Matear, R., Meissner, K., and England, M.: Southern Hemisphere westerlies as a driver of the early deglacial atmospheric CO2 rise, Nat. Commun., 9, 2503,, 2018. a, b

Menviel, L., Capron, E., Tarasov, L., and Abe-Ouchi, A.: The penultimate deglaciation: protocol for PMIP4 transient numerical simulations between 140 and 127 ka, version 1.0, Australian National Data Service,, 2019. a

Mokeddem, Z. and McManus, J.: Persistent climatic and oceanographic oscillations in the subpolar North Atlantic during the MIS 6 glaciation and MIS 5 interglacial, Paleoceanography, 31,, 2016. a, b, c

Mokeddem, Z., McManus, J., and Oppo, D.: Oceanographic dynamics and the end of the last interglacial in the subpolar North Atlantic, P. Natl. Acad. Sci. USA, 111, 11263–11268,, 2014. a

Montade, V., Kageyama, M., Combourieu-Nebout, N., Ledru, M.-P., Michel, E., Siani, G., and Kissel, C.: Teleconnection between the Intertropical Convergence Zone and southern westerly winds throughout the last deglaciation, Geology, 43, 735–738,, 2015. a

Mudelsee, M.: Ramp function regression: a tool for quantifying climate transitions, Comput. Geosci., 26, 293–307,, 2000. a, b, c

Mudelsee, M.: Break function regression, The European Physical Journal Special Topics, 174, 49–63,, 2009. a, b, c

Mulitza, S., Prange, M., Stuut, J.-B., Zabel, M., von Dobeneck, T., Itambi, A. C., Nizou, J., Schulz, M., and Wefer, G.: Sahel megadroughts triggered by glacial slowdowns of Atlantic meridional overturning, Paleoceanography, 23, pA4206,, 2008. a

Murton, J., Bateman, M., Dallimore, S., Teller, J., and Yang, Z.: Identification of Younger Dryas outburst flood path from Lake Agassiz to the Arctic Ocean, Nature, 464, 740–743, 2010. a

Muschitiello, F., Pausata, F., Watson, J., Smittenberg, R., Salih, A., Brooks, S., Whitehouse, N., Karlatou-Charalampopoulou, A., and Wohlfarth, B.: Fennoscandian freshwater control on Greenland hydroclimate shifts at the onset of the Younger Dryas, Nat. Commun., 6, 8939,, 2015. a

NEEM community members: Eemian interglacial reconstructed from a Greenland folded ice core, Nature, 493, 489–494,, 2013. a, b

Ng, H., Robinson, L., McManus, J., Mohamed, K., Jacobel, A., Ivanovic, R., Gregoire, L., and Chen, T.: Coherent deglacial changes in deep Atlantic Ocean circulation, Nat. Commun., 9, 2947,, 2018. a, b

Niedermeyer, E., Prange, M., Mulitza, S., Mollenhauer, G., Schefuss, E., and Schulz, M.: Extratropical forcing of Sahel aridity during Heinrich stadials, Geophys. Res. Lett., 36, L20707,, 2009. a

North Greenland Ice Core project members: High-resolution record of Northern Hemisphere climate extending into the last interglacial period, Nature, 431, 147–151, 2004. a

Okazaki, Y., Timmermann, A., Menviel, L., Harada, N., Abe-Ouchi, A., Chikamoto, M., Mouchet, A., and Asahi, H.: Deep water formation in the North Pacific during the Last Glacial termination, Science, 329, 200–204, 2010. a, b

Oppo, D., McManus, J., and Cullen, J.: Evolution and demise of the Last Interglacial warmth in the subpolar North Atlantic, Quaternary Sci. Rev., 25, 3268–3277, 2006. a, b, c, d, e, f, g, h, i, j

Oppo, D. W., Horowitz, M., and Lehman, S. J.: Marine core evidence for reduced deep water production during Termination II followed by a relatively stable substage 5e (Eemian), Paleoceanography, 12, 51–63,, 1997. a

Otto-Bliesner, B., Russell, J., Clark, P., Liu, Z., Overpeck, J., Konecky, B., deMenocal, P., Nicholson, S., He, F., and Lu, Z.: Coherent changes of southeastern equatorial and northern African rainfall during the last deglaciation, Science, 346, 1223–1227,, 2014. a, b, c

Otto-Bliesner, B. L., Braconnot, P., Harrison, S. P., Lunt, D. J., Abe-Ouchi, A., Albani, S., Bartlein, P. J., Capron, E., Carlson, A. E., Dutton, A., Fischer, H., Goelzer, H., Govin, A., Haywood, A., Joos, F., LeGrande, A. N., Lipscomb, W. H., Lohmann, G., Mahowald, N., Nehrbass-Ahles, C., Pausata, F. S. R., Peterschmitt, J.-Y., Phipps, S. J., Renssen, H., and Zhang, Q.: The PMIP4 contribution to CMIP6 – Part 2: Two interglacials, scientific objective and experimental design for Holocene and Last Interglacial simulations, Geosci. Model Dev., 10, 3979–4003,, 2017. a, b, c, d, e, f

Pahnke, K., Zahn, R., Elderfield, H., and Schulz, M.: 340,000-year centennial-scale marine record of Southern Hemisphere climatic oscillation, Science, 301, 948–952, 2003. a, b

Past Interglacials Working Group of PAGES: Interglacials of the last 800,000 years, Rev. Geophys., 54, 162–219,, 2016. a, b

Pausata, F., Battisti, D., Nisancioglu, K., and Bitz, C.: Chinese stalagmite δ18O controlled by changes in the Indian monsoon during a simulated Heinrich event, Nat. Geosci., 4, 474–480, 2011. a

Payne, A.: A thermomechanical model of ice flow in West Antarctica, Clim. Dynam., 15, 115–125, 1999. a

Pedro, J., Bostock, H., Bitz, C., He, F., Vandergoes, M., Steig, E., Chase, B., Krause, C., Rasmussen, S., Markle, B., and Cortese, G.: The spatial extent and dynamics of the Antarctic Cold Reversal, Nat. Geosci., 9, 51–55,, 2016. a, b

Peltier, W. R. and Drummond, R.: Rheological stratification of the lithosphere: A direct inference based upon the geodetically observed pattern of the glacial isostatic adjustment of the North American continent, Geophys. Res. Lett., 35, L16314,, 2008. a

Peltier, W., Argus, D., and Drummond, R.: Space geodesy constrains ice-age terminal deglaciation: The global ICE-6G-C (VM5a) model, J. Geophys. Res.-Solid Earth, 120, 450–487,, 2015. a

Peterson, L., Haug, G., Hughen, K., and Rohl, U.: Rapid changes in the hydrologic cycle of the Tropical Atlantic during the Last Glacial, Science, 290, 1947–1951, 2000. a

Petit, J., Jouzel, J., Raynaud, D., Barkov, N., Barnola, J.-M., Basile, I., Bender, M., Chappellaz, J., Davis, M., Delaygue, G., Delmotte, M., Kotlyakov, V., Legrand, M., Lipenkov, V., Lorius, C., Pepin, L., Ritz, C., Saltzman, E., and Stievenard, M.: Climate and atmospheric history of the past 420,000 years from the Vostok ice core, Antarctica, Nature, 399, 429–436, 1999. a

Pollard, D. and DeConto, R. M.: Description of a hybrid ice sheet-shelf model, and application to Antarctica, Geosci. Model Dev., 5, 1273–1295,, 2012. a, b

Rabineau, M., Berné, S., Olivet, J.-L., Aslanian, D., Guillocheau, F., and Joseph, P.: Paleo sea levels reconsidered from direct observation of paleoshoreline position during Glacial Maxima (for the last 500,000 yr), Earth Planet. Sci. Lett., 252, 119–137,, 2006. a

Raymo, M. E., Oppo, D. W., Flower, B. P., Hodell, D. A., McManus, J. F., Venz, K. A., Kleiven, K. F., and McIntyre, K.: Stability of North Atlantic water masses in face of pronounced climate variability during the Pleistocene, Paleoceanography, 19, pA2008,, 2004. a

Renssen, H., van Geel, B., van der Plicht, J., and Magny, M.: Reduced solar activity as a trigger for the start of the Younger Dryas?, Quaternary Int., 68–71, 373–383,, 2000. a

Renssen, H., Mairesse, A., Goosse, H., Mathiot, P., Heiri, O., Roche, D., Nisancioglu, K., and Valdes, P.: Multiple causes of the Younger Dryas cold period, Nat. Geosci., 8, 946–949, 2015. a

Risebrobakken, B., Balbon, E., Dokken, T., Jansen, E., Kissel, C., Labeyrie, L., Richter, T., and Senneset, L.: The penultimate deglaciation: high-resolution paleoceanographic evidence from a north-south transect along the eastern Nordic Seas, Earth Planet. Sci. Lett., 241, 505–516,, 2006. a, b

Risebrobakken, B., Dokken, T., Ottera, O., Jansen, E., Gao, Y., and Drange, H.: Inception of the Northern European ice sheet due to contrasting ocean and insolation forcing, Quaternary Res., 67, 128–135, 2007. a

Ritz, S., Stocker, T., Grimalt, J., Menviel, L., and Timmermann, A.: Estimated strength of the Atlantic overturning circulation during the last deglaciation, Nat. Geosci., 6, 208–212, 2013. a

Roberts, N., Piotrowski, A., McManus, J., and Keigwin, L.: Synchronous Deglacial Overturning and Water Mass Source Changes, Science, 327, 75–78,, 2010. a

Roberts, W. H. G. and Valdes, P. J.: Green Mountains and White Plains: The Effect of Northern Hemisphere Ice Sheets on the Global Energy Budget, J. Climate, 30, 3887–3905,, 2017. a

Roche, D. M., Renssen, H., Paillard, D., and Levavasseur, G.: Deciphering the spatio-temporal complexity of climate change of the last deglaciation: a model analysis, Clim. Past, 7, 591–602,, 2011. a, b, c

Rodrigues, T., Alonso-García, M., Hodell, D., Rufino, M., Naughton, F., Grimalt, J., Voelker, A., and Abrantes, F.: A 1-Ma record of sea surface temperature and extreme cooling events in the North Atlantic: A perspective from the Iberian Margin, Quaternary Sci. Rev., 172, 118–130,, 2017. a, b

Rohling, E., Grant, K., Bolshaw, M., Roberts, A. P., Siddall, M., Hemleben, C., and Kucera, M.: Antarctic temperature and global sea level closely coupled over the past five glacial cycles, Nat. Geosci., 2, 500–504, 2009. a

Rohling, E. J., Hibbert, F. D., Williams, F. H., Grant, K. M., Marino, G., Foster, G. L., Hennekam, R., de Lange, G. J., Roberts, A. P., Yu, J., Webster, J. M., and Yokoyama, Y.: Differences between the last two glacial maxima and implications for ice-sheet, δ18O, and sea-level reconstructions, Quaternary Sci. Rev., 176, 1–28,, 2017. a, b, c, d, e

Rosell-Melé, A. and Prahl, F. G.: Seasonality of UK'37 temperature estimates as inferred from sediment trap data, Quaternary Sci. Rev., 72, 128–136,, 2013. a

Schilt, A., Baumgartner, M., Blunier, T., Schwander, J., Spahni, R., Fischer, H., and Stocker, T.: Glacial-interglacial and millennial-scale variations in the atmospheric nitrous oxide concentration during the last 800,000 years, Quaternary Sci. Rev., 29, 182–192,, 2010. a, b, c, d, e

Schmittner, A. and Lund, D. C.: Early deglacial Atlantic overturning decline and its role in atmospheric CO2 rise inferred from carbon isotopes (δ13C), Clim. Past, 11, 135–152,, 2015. a

Schmittner, A., Bostock, H., Cartapanis, O., Curry, W., Filipsson, H., Galbraith, E., Gottschalk, J., Herguera, J., Hoogakker, B., Jaccard, S., Lisiecki, L., Lund, D., Martínez-Méndez, G., Lynch-Stieglitz, J., Mackensen, A., Michel, E., Mix, A., Oppo, D., Peterson, C., Repschläger, J., Sikes, E., Spero, H., and Waelbroeck, C.: Calibration of the carbon isotope composition (δ13C) of benthic foraminifera, Paleoceanography, 32, 512–530,, 2017. a

Schneider, R., Schmitt, J., Köhler, P., Joos, F., and Fischer, H.: A reconstruction of atmospheric carbon dioxide and its stable carbon isotopic composition from the penultimate glacial maximum to the last glacial inception, Clim. Past, 9, 2507–2523,, 2013. a, b, c, d, e, f, g

Shackleton, N.: Les methodes quantitatives d'etude des variations du climat au cours du Pleistocene, vol. 219, 203–210, CNRS, 1974. a

Shackleton, N., Sanchez-Goni, M., Pailler, D., and Lancelot, Y.: Marine Isotope Substage 5e and the Eemian Interglacial, Global Planet. Change, 36, 151–155,, 2003. a, b, c, d

Shakun, J., Clark, P., He, F., Marcott, S., Mix, A., Liu, Z., Otto-Bliesner, B., Schmittner, A., and Bard, E.: Global warming preceded by carbon dioxide concentrations during the last deglaciation, Nature, 484, 49–55, 2012. a, b

Siddall, M., Rohling, E. J., Almogi-Labin, A., Hemleben, C., Meischner, D., Schmelzer, I., and Smeed, D. A.: Sea-level fluctuations during the last glacial cycle, Nature, 423, 853–858, 2003. a

Siddall, M., Bard, E., Rohling, E., and Hemleben, C.: Sea-level reversal during Termination II, Geology, 34, 817,, 2006. a

Sigman, D., Hain, M., and Haug, G.: The polar ocean and glacial cycles in atmospheric CO2 concentration, Nature, 466, 47–55,, 2010. a

Sime, L., Wolff, E., Oliver, K., and Tindall, J.: Evidence for warmer interglacials in East Antarctic ice cores, Nature, 462, 342–345,, 2009. a

Skinner, L. and Shackleton, N.: Deconstructing Terminations I and II: revisiting the glacioeustatic paradigm based on deep-water temperature estimates, Quaternary Sci. Rev., 25, 3312–3321, 2006. a

Skinner, L., Fallon, S., Waelbroeck, C., Michel, E., and Barker, S.: Ventilation of the deep Southern Ocean and deglacial CO2 rise, Science, 328, 1147–1151, 2010. a

Spahni, R., Chappellaz, J., Stocker, T., Loulergue, L., Hausammann, G., Kawamura, K., Flückiger, J., Schwander, J., Raynaud, D., Masson-Delmotte, V., and Jouzel, J.: Atmospheric methane and nitrous oxide of the late Pleistocene from Antarctic ice cores, Science, 310, 1317–1321, 2005. a

Stein, M., Wasserburg, G., Aharon, P., Chen, J., Zhu, Z., Bloom, A., and Chappell, J.: TIMS U-series dating and stable isotopes of the last interglacial event in Papua New Guinea, Geochim. Cosmochim. Acta, 57, 2541–2554,, 1993. a

Stephens, B. and Keeling, R.: The influence of Antarctic sea ice on glacial-interglacial CO2 variations, Nature, 404, 171–174, 2000. a

Stirling, C., Esat, T., Lambeck, K., and Culloch, M.: Timing and duration of the Last Interglacial: evidence for a restricted interval of widespread coral reef growth, Earth Planet. Sci. Lett., 160, 745–762, 1998. a, b, c

Stocker, T. and Johnsen, S.: A minimum thermodynamic model for the bipolar seesaw, Paleoceanography, 18, 1087,, 2003. a

Stone, E. J., Capron, E., Lunt, D. J., Payne, A. J., Singarayer, J. S., Valdes, P. J., and Wolff, E. W.: Impact of meltwater on high-latitude early Last Interglacial climate, Climate of the Past, 12, 1919–1932,, 2016. a, b

Stouffer, R., Seidov, D., and Haupt, B.: Climate response to external sources of freshwater: North Atlantic versus the Southern Ocean, J. Climate, 20, 436–448, 2007. a, b, c, d

Svendsen, J., Alexanderson, H., Astakhov, V., Demidov, I., Dowdeswell, J., Funder, S., Gataullin, V., Henriksen, M., Hjort, C., Houmark-Nielsen, M., Hubberten, H., Ingolfsson, O., Jakobsson, M., Kjaer, K., Larsen, E., Lokrantz, H., J. Pekka Lunkkap, A. L., Mangerud, J., Matiouchkov, A., Murray, A., Möller, P., Niessen, F., Nikolskaya, O., Polyak, L., Saarnisto, M., Siegert, C., Siegert, M., Spielhagen, R., and Stein, R.: Late Quaternary ice sheet history of northern Eurasia, Quaternary Sci. Rev., 23, 1229–1271, 2004. a, b, c

Swingedouw, D., Mignot, J., Braconnot, P., Mosquet, E., Kageyama, M., and Alkama, R.: Impact of Freshwater Release in the North Atlantic under Different Climate Conditions in an OAGCM, J. Climate, 22, 6377–6403,, 2009. a

Syverson, K. and Colgan, P.: Quaternary Glaciations: Extent and Chronology – A Closer Look, vol. 15 of Developments in Quaternary Science, chap. The Quaternary of Wisconsin: An updated review of stratigraphy, glacial history and landforms, 537–552, Elsevier, 2011. a, b

Tarasov, L.: The global GLAC-1c deglaciation chronology, melwater pulse 1-a, and a question of missing ice, IGS Symposium: Contribution of Glaciers and Ice Sheets to Sea-Level Change, Chamonix, France, 26–30 May, 2014. a

Tarasov, L. and Peltier, W. R.: Terminating the 100 kyr Ice Age cycle, J. Geophys. Res., 102, 21665–21693, 1997. a

Tarasov, L. and Peltier, W.: Arctic freshwater forcing of the Younger Dryas cold reversal, Nature, 435, 662–665, 2005. a

Tarasov, L. and Peltier, W.: A calibrated deglacial drainage chronology for the North American continent: evidence of an Arctic trigger for the Younger Dryas, Quaternary Sci. Rev., 25, 659–688, 2006. a

Tarasov, L. and Peltier, W. R.: The Co-evolution of continental ice cover and permafrost extent over the last glacial-interglacial cycle in North America, J. Geophys. Res., 112, F02S08,, 2007. a

Tarasov, L., Dyke, A. S., Neal, R. M., and Peltier, W. R.: A data-calibrated distribution of deglacial chronologies for the North American ice complex from glaciological modeling, Earth Planet. Sci. Lett., 315–316, 30–40, 2012. a, b, c, d, e, f, g, h

Taylor, K. E., Stouffer, R. J., and Meehl, G. A.: An Overview of CMIP5 and the Experiment Design, B. Am. Meteorol. Soc., 93, 485–498,, 2012. a

Thomas, A. L., Henderson, G. M., Deschamps, P., Yokoyama, Y., Mason, A. J., Bard, E., Hamelin, B., Durand, N., and Camoin, G.: Penultimate Deglacial Sea-Level Timing from Uranium/Thorium Dating of Tahitian Corals, Science, 324, 1186–1189,, 2009. a, b, c, d

Thornalley, D., Barker, S., Broecker, W., Elderfield, H., and McCave, I.: The deglacial evolution of North Atlantic Deep Convection, Science, 331, 202–205, 2011. a

Timm, O., Köhler, P., Timmermann, A., and Menviel, L.: Mechanisms for the onset of the African Humid Period and Sahara Greening 14.5–11 ka BP, J. Climate, 23, 2612–2633, 2010. a

Timmermann, A., An, S. I., Krebs, U., and Goosse, H.: ENSO suppression due to a weakening of the North Atlantic thermohaline circulation, J. Climate, 18, 3122–3139, 2005. a

Toggweiler, J. and Lea, D.: Temperature differences between the hemispheres and ice age climate variability, Paleoceanography, 25, F02S08,, 2010. a

Toggweiler, J., Russell, J., and Carson, S.: Midlatitude westerlies, atmospheric CO2, and climate change during ice ages, Paleoceanography, 21, PA2005,, 2006. a

Tschumi, J. and Stauffer, B.: Reconstructing past atmospheric CO2 concentration based on ice-core analyses: open questions due to in situ production of CO2 in the ice, J. Glaciol., 46, 45–53, 2000. a

Tzedakis, P., Frogley, M., and Heaton, T.: Last Interglacial conditions in southern Europe: evidence from Ioannina, northwest Greece, Global Planet. Change, 36, 157–170, 2003. a, b

Tzedakis, P., Crucifix, M., Mitsui, T., and Wolff, E.: A simple rule to determine which insolation cycles lead to interglacials, Nature, 542, 427–432, 2017. a

Tzedakis, P., Drysdale, R., Margari, V., Skinner, L., Menviel, L., Rhodes, R. H., Taschetto, A. S., Hodell, D. A., Crowhurst, S. J., Hellstrom, J. C., Fallick, A. E., Grimalt, J. O., McManus, J. F., Martrat, B., Mokeddem, Z., Parrenin, F., Regattieri, E., Roe, K., and Zanchetta, G.: Enhanced climate instability in the North Atlantic and southern Europe during the Last Interglacial, Nat. Commun., 9, 4235, 2018. a, b, c, d

Tzedakis, P. C., Wolff, E. W., Skinner, L. C., Brovkin, V., Hodell, D. A., McManus, J. F., and Raynaud, D.: Can we predict the duration of an interglacial?, Clim. Past, 8, 1473–1485,, 2012. a

Tzedakis, P. C., Emerson, B., and Hewitt, G.: Cryptic or mystic? Glacial tree refugia in northern Europe, Trend. Ecol. Evolut., 28, 696–704,, 2013. a

Uemura, R., Masson-Delmotte, V., Jouzel, J., Landais, A., Motoyama, H., and Stenni, B.: Ranges of moisture-source temperature estimated from Antarctic ice cores stable isotope records over glacial-interglacial cycles, Clim. Past, 8, 1109–1125,, 2012. a

Uemura, R., Motoyama, H., Masson-Delmotte, V., Jouzel, J., Kawamura, K., Goto-Azuma, K., Fujita, S., Kuramoto, T., Hirabayashi, M., Miyake, T., Ohno, H., Fujita, K., Abe-Ouchi, A., Iizuka, Y., Horikawa, S., Igarashi, M., Suzuki, K., Suzuki, T., and Fujii, Y.: Asynchrony between Antarctic temperature and CO2 associated with obliquity over the past 720,000 years, Nat. Commun., 9, 961,, 2018. a, b

van Kreveld, S., Samthein, M., Erlenkeuser, H., Grootes, P., Jung, S., Nadeau, M., Pflaumann, U., and Voelker, A.: Potential links between surging ice sheets, circulation changes, and the Dansgaard-Oeschger cycles in the Irminger Sea, 60–18 kyr, Paleoceanography, 15, 425–442, 2000. a, b

Vázquez Riveiros, N., Waelbroeck, C., Skinner, L., Duplessy, J.-C., McManus, J. F., Kandiano, E. S., and Bauch, H. A.: The ”MIS 11 paradox'” and ocean circulation: Role of millennial scale events, Earth Planet. Sci. Lett., 371, 258–268,, 2013. a, b

Veres, D., Bazin, L., Landais, A., Toyé Mahamadou Kele, H., Lemieux-Dudon, B., Parrenin, F., Martinerie, P., Blayo, E., Blunier, T., Capron, E., Chappellaz, J., Rasmussen, S. O., Severi, M., Svensson, A., Vinther, B., and Wolff, E. W.: The Antarctic ice core chronology (AICC2012): an optimized multi-parameter and multi-site dating approach for the last 120 thousand years, Clim. Past, 9, 1733–1748,, 2013. a, b, c, d, e, f

Wang, Y., Cheng, H., Edwards, R., An, Z., Wu, J., Shen, C., and Dorale, J.: A high-resolution absolute-dated Late Pleistocene monsoon record from Hulu Cave, China, Science, 294, 2345–2348, 2001. a

Weaver, A., Saenko, O., Clark, P., and Mitrovica, J.: Meltwater pulse 1A from Antarctica as a trigger of the Bølling-Allerød warm interval, Science, 299, 1709–1713, 2003. a

Weber, M., Clark, P., Kuhn, G., Timmermann, A., Sprenk, D., Gladstone, R., Zhang, X., Lohmann, G., Menviel, L., Chikamoto, M., Friedrich, T., and Ohlwein, C.: Millennial-scale variability in Antarctic ice-sheet discharge during the last deglaciation, Nature, 510, 134–138,, 2014. a

Wilson, G. P., Reed, J. M., Frogley, M. R., Hughes, P. D., and Tzedakis, P. C.: Reconciling diverse lacustrine and terrestrial system response to penultimate deglacial warming in southern Europe, Geology, 43, 819,, 2015. a

Yang, S. and Ding, Z.: A 249 kyr stack of eight loess grain size records from northern China documenting millennial-scale climate variability, Geochem. Geophys. Geosyst., 15, 798–814, 2014. a, b, c, d

Yau, A., Bender, M., Robinson, A., and Brook, E.: Reconstructing the last interglacial at Summit, Greenland: Insights from GISP2, P. Natl. Acad. Sci. USA, 113, 9710–9715,, 2016.  a

Zhang, X., Lohmann, G., Knorr, G., and Xu, X.: Different ocean states and transient characteristics in Last Glacial Maximum simulations and implications for deglaciation, Clim. Past, 9, 2319–2333,, 2013. a, b

Zhang, X., Lohmann, G., Knorr, G., and Purcell, C.: Abrupt glacial climate shifts controlled by ice sheet changes, Nature, 512, 290–294,, 2014. a, b

Zhu, J., Liu, Z., Brady, E., Otto-Bliesner, B., Zhang, J., Noone, D., Tomas, R., Nusbaumer, J., Wong, T., Jahn, A., and Tabor, C.: Reduced ENSO variability at the LGM revealed by an isotope-enabled Earth system model, Geophys. Res. Lett., 44, 6984–6992,, 2017. a

Publications Copernicus
Short summary
As part of the Past Global Changes (PAGES) working group on Quaternary Interglacials, we propose a protocol to perform transient simulations of the penultimate deglaciation for the Paleoclimate Modelling Intercomparison Project (PMIP4). This design includes time-varying changes in orbital forcing, greenhouse gas concentrations, continental ice sheets as well as freshwater input from the disintegration of continental ice sheets. Key paleo-records for model-data comparison are also included.
As part of the Past Global Changes (PAGES) working group on Quaternary Interglacials, we propose...